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SMITHSONIAN INSTITUTION
UNITED STATES NATIONAL MUSEUM
ButteTin 161
THE FORAMINIFERA OF THE
TROPICAL PACIFIC COLLECTIONS OF
THE “ALBATROSS,” 1899-1900
Part 1.—ASTRORHIZIDAE to TROCHAMMINIDAE
BY
JOSEPH AUGUSTINE CUSHMAN
Director, Cushman Laboratory for Foraminiferal Research
UNITED STATES
GOVERNMENT PRINTING OFFICE
WASHINGTON : 1932
ADVERTISEMENT
The scientific publications of the National Museum include two
series, known, respectively, as Proceedings and Bulletin.
The Proceedings series, begun in 1878, is intended primarily as a
medium for the publication of original papers, based on the collec-
tions of the National Museum, that set forth newly acquired facts
in biology, anthropology, and geology, with descriptions of new
forms and revisions of limited groups. Copies of each paper, in
pamphlet form, are distributed as published to libraries and scientific
organizations and to specialists and others interested in the different
subjects.
The dates at which these separate papers are published are re-
corded in the table of contents of each of the volumes.
The Bulletin series, the first of which was issued in 1875, con-
sists of separate publications comprising monographs of large zoo-
/logieal groups and “other, general systematic treatises (occasionally
on
‘fin several volumes), faunal works, reports of expeditions, cata-
. logues of type specimens and special collections, and other material of
similar nature. The majority of the volumes are octavo in size,
. f
but’a’ quarto sjze-ha& heen adopted in a few instances in which large
plates*were-regarded as indispensable. In the Bulletin series appear
volumes under the heading Contributions from the United States
National Herbarium, in octavo form, published by the National
Museum since 1902, which contain papers relating to the botanical
collections of the Museum.
The present volume forms No. 161 of the Bulletin series.
ALEXANDER WETMORE,
Assistant Secretary, Smithsonian Institution.
Wasuineron, D. C., May 12, 1932.
II
CONTENTS
Page
BPEL TOT ME IN oe ep Steele Ot ec nas AM 1
PRAM A SIOPMIZIARC aoa cece OOO 4
Genus habdamming sen cecicen ature eaten 4
GUYSIORUM sam Liane oe nes ee ee 4
Bamily, Reophacidae..<oqunese cee ee et ER ke oe 4
IS CTTUB VICE ONES Serene reer tee Re Cue MEI oo ie 4
OUGUGUMNAUUS oat ese ate ec I AS eee) 4
Bamitly Wo WuONaae ee rea el oc - 5
GenusrHaplophragmotdes= ~~ 22 eee) __ 5
COUPES ALA = Leer eee ee, oS 5
Bamily AMMO disCldses = 2 ree eee eee a OMG 5
Genus: Ammodtsciels ac on eee eee eee AR 5
TOC TEND yet etre eh yeaa tee gs ak a I 5
bcrarl WeePeXUUIATHO RG. ah ee ee tee ee OR 6
Subfamily Spiroplectammininae..--—-.<22 2-2-2222. Se 6
GeausiSpiroplectamming .. 2... =... ce ot ooioe? pyre 6
TUE EASE rater ne ented Ca 7
Sublamily. Pextularinae =e ee ek 7
COUN ELLA RE A do se ee ene ee OD IRE 74
OLVRECE eRe ep re eee ey SE 8 ic 8
VEE WOCCERTED 225 te S wn e S R 8
SOMGIGLGL Dt REE AE. le eee NE lp 9
LLL CLS SOR oe OD PORE 09 8 Ee Sy OR Pe 9
LTTE LS SAS Ce ee 1s 10
COTE TOR SE eB ee 2 BAAS DOMNDE LTA Lu berm, 11
COMMUG UGE Sa take ie eee ee eh ee 12
LER O SSE Re ee SON ose fap ae RNG 12
CONCOUG Se nm tre ee er oe ee ee 13
Pua yak V EEN UATE ACD 2 as ale ee hi ee wy ee a epee 13
Genus Gmudnyimatecccs cece eee tein SE 13
DU Bis pce pc enn NR he I LI 13
intqngulares Vari angulata. 22.2 ae eie 2 SS 14
QUOUTONPULET San ceie wetiiee nn eReweee eel eh... DUANE ot Ly 14
OST) ee a ee ee eee een SC | kee 15
BUG OS Oe or ete eee ee ae, SE © 15
RSI SVC TQUUL Ne: = fe le cre ne Ee ie oe 16
COMMUNE 5 a ee DE 16
BECHLER br Stee eR ee eee RL, Oe De 16
DOT NEE feet ee ese eee ee ee) ie 20 17
MeSeET Ty gv ENON AOU SORA ee ae ht re rei ee ee ee RE Ly
MGeETAUIS IVETE UCLOCUININE pectoral ee NO 17
anguina:varliagglutinans « «=< cn. SOLA. 18
BTR TC CLO ER ER hr teh Heart RN SUS 19
BERET OLOO SG ORs eee eee ie OUT AED 20
OSU a Hearne Are re tp I 2 20
PRES WAT ASUDeUNneOG sek ete SEO 21
CTC SBR Ga oo anesthe ee ar ra ed 21
FUME] ULEOTIBES Sanpete corer I NY TB 22
IV BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Family Miliolidae—Continued.
Genus Quinqueloculina—Continued. Page
ef; gractlis... 2 SAA See hs ee Eo es eee 23
OPUBCLULL ONLINE a gS a Es at a 23
SAA Pad gl ola 0H (Hs Pala ten ee LUA LS A le ALOU) Pa NRE ty A 24
PHOTOG ee i Os a a i eae ee 25
nolygona. 22> eke ees eee ae. ee eee 25
SQMOGENSIS 2222 e eek De a es OE eee 2 26
SCMITCLICULOSAS Les a SNe es oe ee pea plaka Wp ie 27
SETURL Cees lS Se UN aA a ag alg 7,
SULCOEG ate le a eae ines ee _ 28
Genus Schlumbergerina22 2-2 282 ee eee ieee ee 29
alveoliniformts... 22-02 22 Le, SUS Ba Ee a eS) 29
GeniisMassalenay 25 lo SR Se 2 eS TR ae el A ae ae VE 30
alveolanifor mis: <= 22 2 ake ahs Ns wR yA ciple OB 30
PROROLG 32 oo eS eee ee a eee ee eee a a 31
{NOCQUGIS 2 ne ee a ee Eee ee 32
USER GUGS ae SR Se I Se eR 32
MACON i 8 se VO oe et SEN OT Sas ari aol ah 33
CTENDGL ewe oe Se Se Sea reg lo ad el ES et oe
Genus Smiroloculing. 2-22-21. 2 lee tae ee bee 34
QR GLCL OUD Sa ee le sage 34
VETS. SCTVUIGIG 25 Ep PTY eee rd ay 35
VAT GCOSCOLG Sa 2 Se Ns ee a a oe a iA BLO 35
antilanum hes = POL a Sn De Ss he a eee eae 36
Weer? cangildta sos oe Fe ahs meme) peg als Sh ie 37
VAT \GOQ Udine See a a La aie ele 2 38
UAT POUT CO SOLS eee ea 0) Ss ik at LU TY 2 | an i a 38
CULL se alas genes oe a one 39
plantssima@ var. samogensis... 223 5522s) Ane See 39
CLEGG feels 2 Se SUG aN BE a ob a Sh ae TN Ea 39
CL whe Ld Se a Se I Re dey tal 40
EL or oe i i a Oh a 41
SD) one ee ee es ea alga 42
Genus Hauerina 23 oo 2 2k BE ERS * a ee eee = 42
fragtlissima. 22202522. eee ee eee ee 42
OTNALTSST NG ao oa a es a el tpt ag age ge 43
MRT nt Ng lea gh 44
Boracay So is I a el a 44
Genus Sigmowlinag c= 3 2s eee ee eee ele Ee 45
Cdwardsts one A | AE ain Ra yopepe pe 45
Genus “Aritculinases fee 2 ee Leap aaetia 46
UTC ok es Es Se Cet dT 47
suloata(R))e ok s Gg 2 2 pe oe 47
«Genus Nubeculina 2. = 3 a he Ae ae ehh ie a ee 48
@waricoia 2. oo ee a a gece phe aN ery gaa 48
VAT. CUCU a aie an oe a lt el at wg LO 49
CRODMENG Ss Se Se ee LL eA 2 apa Age AL ap 49
Genus: Priloculina s = Ose als a Nae a aA peal eR. DeLee 50
ODL ON GE 2 a 2s RT TO et a 2 a er 50
CONCULETES 202 ooh ho Se i eal aR al a aise a re 52
Lem 8 ao es ica a a mV a 53
VE. SDGTSTCOSEGLGS oy LW SOE oA Gale Dee ety yee 54
CONTENTS Wy:
Family Miliolidae—Continued.
Genus Triloculina—Continued. Page
DIEPEMULIT IRS ee ee Rae ee. 2 oh he Nd a, 54
CO CECMUL CC ae nate ae ee ae ee ae Nene ee ete ee St ee S 54
ELTA UL TULL LT) scat ce eee ere eS ee ok 55
entelLIGne SS SOLO RS foe a eee eae ee ee rk SS 55
INU TUCLE Ra eet a pn Sie Ns ea ea ete eI il Bis Spd 56
RETOLD ee SA eR ATES, 5 o/h SR Ne EN 2 56
URES EM EC Cte a co ee ahh ay ne eR en 57
TET T Se oh eee seco aaa th TE ae Ee eee a a, Nae 58
EN TCOT TTC QUC See Noe als ie real NN EE SS eee aes SS 59
DET RELATUGIeG a el tS a ok a UUs Ein ab eA paiena bm Si 60
LEO E IDES ee lS ek ae eet ee oer ey ees ets Es 60
Space Ce ea Ee LEE) eee Bp ed Nei s 61
ROTM EE Oe ad LS RUN RN SEN RI eye RN Fe Se 61
CLETGLUCULUCLLC, es nue ser Saki <tc Vea ENA, ne ar EM go 2 62
SVGUTEs SUNG OLE ee ea hs Ae es epee ae = er ie aa 63
AUTEN PRUNE CL See ol pe ee ype RS ER yt a A get Se 64
globule: ey a ee RE Ne ie Ge ee A 65
TELTIC CURE ere ne nae Ee RT RY SENN OS ee oe 66
fame Ophnthalmidiudae ss). Jeo sora see Oe ULE le Eee 67
Sapramity)Comuspirinae.- us Steet we Poe ee ek ees oe 67
Genus Gomis pias ss Wigs ly VE lin ls es brie celal d 67
ARVO EIS Sinne Seen ie el pra Ale ec RR ea Ps 67
Subfamily. Nodobaculariinae..- =... 2 _ estes sto eomgere _ 5 68
Genus Nodobacularia= .'—- 2aetaes eS Bee pane tin eainnne 5 68
Ontilanumevar PacificaeTs ys ok Weis peewee ea eel ae 68
ATUL LUGE Eas Ow pee Tick ey loa | pili alten di a at at aa ages ols fat LI 69
Subfamily ;Ophthalmidiinae_ 2 <iseuds. sop see ees ode Sa eee 70
Genus. Spironthalmidiums. 2 322k eee rosy iy ay ae 70
acutimangos._ 2 = Peewee ee nos ene. I poten fF 70
GenusiPlanis piri were. Au von Leman Baie ot 71
CLULGU GREE ee ae EY ann noe DE See hey teh ake 71
GURICHIOLGL he Eine BR tn i a pet ap ag are 2 72
GenusiwWertebralintaisss. So eee ee) oie nl anand se 235 eal d 72
SUMOIG 22 ee Ces rae men = SE a re econ sens ee 73
Subfamily Nubeculariinge: sisse!s sue te eh Po Sains 74
GenustParrenaes) CS. Ue! EE BO Sa Eo Te 74
RCL ee CR od et eB ee NS ee ea ere 74
Ramilyebischeninid tes, Sete ge ss ee ee a ed ae See 75
SERGIARES WE ROCHA a a ce hak AE hel gs 75
DOU TUCUN Othe wie mils eo ns ieee ee eres ee ae ee 75
[A Rech Papen arin kA Nea es A MN a We ak ge Rg LI SE OR la 76
ENVOLUEE ae) RIES COREE ATS Sy EERE EY AT ae 76
iam SP och amarnini dae t's yyy sa NS yd oy ee aa ee pen Dee 77
BSrplAmiily erOchamiIMininne 2) oc 8 bs ye en ee Ss ee 77
Gens, Brockammingsn 2 oo ok oe on ee Bee, 77
BIO UIVA LO eee ee ce es Re) eS Ste NS er a Re ap Ses 77
SubtamilyGlopotextularimae 202 ob!) es ee a ore Cl
COTM OUPER Tae IO LN ok Dok CE Re ae re ek a Ue ae
PolmInarnnnotdess 1 Rie ott. Sh hs een eer 78
Pisin A IOs AD IALes eS See ee as ye oe ee ge oe 79
ILLUSTRATIONS
TEXT FIGURES
Page
Ficure 1. General route of the Albatross, 1899-1900_________________- 3
PLATES
Facing page
PuaTE 1. Species of Reophax, Haplophragmoides, Spiroplectammina, and
cD eactanbenr ies seh i bn ea Ng 84
2.. Species: of Tecwularie.... 8... 2... ae eae ae eae 84
3.. Species-of -Tertularta.and..Gaudryina_.__ sau =4ehe, eee e lh | 84
4. Species of Gaudryina and Clavulina__----------------------- 84
5-7... Species of Quinguelocultna.: 0.25. 22).--2 25 ee 84
8. Species of Schlumbergerina, Massilina, and Spiroloculina__-__-_-- 84
9... Species:of. Spiroloculmasveso.... .- sucitdeeln dn, eee 84
10. Species of Spiroloculina and Hauerina___---.---------------- 84
11. Species of Articulina, Nubeculina, Sigmoilina, and Triloculina_-_ 84
12) 18. Speciesot Triloculiag. .=. 22 4223 ee ee eee 84
14,/16.. Spegies Gh Pyrgo.ae oo eee cee ne eke eee eee 84
16. Species of Nodobacularia, Cornuspira, Spiropthalmidium, Plani- 84
spirenajand Vertebraling.. 222225. Uses seco eee eee 84
17. Species of Parrina, Fischerina, Trochammina, and Nouria------ 84
vi
li
THE FORAMINIFERA OF THE TROPICAL PACIFIC
COLLECTIONS OF THE “ALBATROSS,” 1899-1900
Part 1.—ASTRORHIZIDAE TO TROCHAMMINIDAE
By JosrrH AUGUSTINE CUSHMAN
Director, Cushman Laboratory for Foraminiferal Research
Sharon, Massachusetts
INTRODUCTION
THIS PAPER is the first part of a work the intent of which is to
describe and illustrate the foraminifera of the tropical Pacific
collected by the United States Bureau of Fisheries steamer Albatross,
together with certain other related material from shallow water of
the same region. Parts 2 and 3 will take up in systematic order
the rest of the families after the Trochamminidae.
Numerous papers deal with the shallow-water foraminifera of the
Indo-Pacific, but from deep water few records are available for the
general area except those from the Challenger expedition of 50
years ago.
Two previous bulletins of the United States National Museum,
one on the foraminifera of the North Pacific Ocean (Bulletin 71,
1910-1917) and the other on the foraminifera of the Atlantic Ocean
(Bulletin 104, 1918-1931), are general accounts, especially of the
foraminifera dredged by the Albatross. These bulletins also include
many of the species recorded from the same areas in other pub-
lications.
The collections of the Albatross made during the voyage from
August 26, 1899, to February 21, 1900, form the basis of the present
series. These were made largely in deep water, although many of
them were in the immediate neighborhood of oceanic islands, par-
ticularly the Paumotu group. Shallow-water collections made from
this same area have also come into my hands for study and greatly
supplement the fauna obtained in the deeper-water dredgings. A
few of the stations are in north latitude, especially those near the
Marshall and Ladrone Islands. The fauna treated, however, is all
more or less of a unit, and the fact that some forms come from north
of the Equator and most of them from south does not interfere with
the general unity of the paper.
a
2 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Most of the papers published on the foraminifera of the Indo-
Pacific region are cited in the synonymy under the various species,
and in some of them species were figured and described that were
not obtained in our collections. To have added all the other records
from the general Indo-Pacific region would not only have greatly
enlarged this work, but would have involved the addition of many
species the types of which were not available for study. It has
seemed best, therefore, to limit the scope of the work to the actual
collections of the Albatross and to the other available shallow-water
deposits from the same groups of islands.
Though the fauna as a whole is a unified one, nevertheless the
study of the present collections shows that there are in the shallow
water about various groups of oceanic islands of the South Pacific
locally developed species that are strictly limited. The species of
deeper water, however, have a wide distribution.
Furthermore, there are numerous species that occur definitely asso-
ciated in other areas. For instance, many are widely distributed
among the islands of the South Pacific that are known also from the
West Indian region. Others are known from the Mediterranean and
the Red Sea regions. A few of them are very closely related to
species originally described from the warm-water facies of the
Miocene or Pliocene of southern Europe.
The accompanying map (fig. 1) shows the general path of the
Albatross during this voyage, and for further details the reader is
referred to the tables accompanying a number of the species. These
give the depth, temperature, and other data regarding the samples
from which the material was taken.
The plates are all from original drawings made from the collec-
tions by Miss Margaret S. Moore, and form perhaps the most useful
part of the present work.
I wish to take this opportunity to express my great appreciation
of the many kindnesses and the constant help that the United States
National Museum and its staff have so constantly given me in the
study of this and other material.
TABLE 1.—Abbreviations used in the tables in this report to denote the char-
acter of the bottoms of the dredging stations
Abbreviation Meaning Abbreviation Meaning Abbreviation Meaning
Di eee eee brown Lisa sate ees light. DUM So ee eee ea pumice
pre Ares. Sars broken Pe. ers eee SE mud. TOY ee eee red
Ca sgh. hbo. Meurer clay. Mang sre. 25 4 manganese: {|| Si2-. 25242. 2.-Liene sand.
OMebe elie Soe ee re ey coral. NOG's2 ess eee nodules. Sse eo ee shells.
Crees ee eee ee coarse. ON ee ee eee ooze. VO ee eee ee volcanic.
fne@siss! cee eS oe fine. Pat! 1222. yee particles. Whr.fisevs eas white.
glob mess ss globigerina. || pter_---.---------- pteropods. Male nee a Re yellow.
BY------------------ gray.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS
OOGI-GGEST ‘880.12907" 9} JO BINoOI [BADIaH—'T TANI]
SANV1ISI1 WONOL -
"S$ 02 NE
SANVISI VOLS" ;
SANVI$1 SYUSANDYYIN ”
yOLYNOZF
SANW'1S1
“N Or os
°
™ OO! “MO! "MOrv!} “MmMO*DlL OB!
4 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Family ASTRORHIZIDAE
Genus RHABDAMMINA M. Sars, 1869
RHABDAMMINA ABYSSORUM M. Sars
There are a few very fragmentary specimens of a tubular form
composed of rather coarse sand grains, firmly cemented. These
strongly resemble broken portions of the arms of this species, and
are placed here provisionally. They are the only records for this
family in the material examined. They are from Albatross Station
H3779, latitude 24° 45’ N., longitude 130° 16’ W., in 2,627 fathoms.
Family REOPHACIDAE
Genus REOPHAX Montfort, 1808
REOPHAX AGGLUTINATUS Cushman
PLATE 1, Figures 1-3
Reophaz agglutinatus CUSHMAN, Proc. U. S. Nat. Mus., vol. 44, p. 637, pl. 79,
fig. 6, 1913; U. S. Nat. Mus. Bull. 104, pt. 2, p. 9, pl. 2, figs. 4, 5, 1920; U. =
Nat. Mus. Bull. 100, vol. 4, p. 73, pl. 14, figs. 2 a, 6, 1921.
Test composed of a series of pyriform chambers with a thin chiti-
nous lining and the exterior composed of tests of other foraminifera
firmly cemented; aperture extended into a narrow, cylindrical neck.
There are in the A/dbatross collections a number of specimens, some
of which are figured here. These are composed of young tests of
nearly uniform size neatly cemented. A few of the specimens simi-
lar to that shown in Plate 1, Figure 1, are evidently the initial cham-
bers of this form. Although the stations are somewhat widely dis-
tant from one another, the specimens are all very uniform in char-
acter. They have been referred to this species, which in other locali-
ties seems to lack the uniformity in the size of the tests that com-
pose the wall.
TABLE 2.—Reophax agglutinatus—material examined
|
Bot-
Num-
Alba- tom
U.S.N.M. | ber of : Character of Occur-
No. | speci- Navel Locality epee ous bottom 1 rence
mens Peres
ture
Fathoms| ° F.
DIGI 2a ES 1 | H3809 | Entrance to Avatoru Pass, 645) 22. 3 fne. wh. co. s.._-| Rare.
Rahiroa Atoll, 2.5 miles S.
21973=2.=--= 1 | H8893 | East point Makemo, 78° N., 11 1, 221 | 26 glob. mang.--.--- Do.
miles W.
21974 === 1 | H3910 | Southwest point Aki Aki, 1 mile 377 | 43 CONG Eee Do.
E.
FGT5 ee 2 2 {Lat. 19° 45’ 30” S., long. 139°
SIGIR S. 2 |rrao19 | 54 W. 1, 494 | 35.4 | glob. mang__-__-_ Do.
1 Key to abbreviations is given in Table 1.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 3
Family LITUOLIDAE
Genus HAPLOPHRAGMOIDES Cushman, 1910
HAPLOPHRAGMOIDES TRULLISSATA (H. B. Brady)
PLATE 1, Fieures 4 a, 0
Trochammina trullissata H. B. Brapy, Quart. Journ. Micr. Sci., vol. 19, p. 56,
pl. 5, figs. 10 a, b, 11, 1879; Rep. Voy. Challenger, Zoology, vol. 9, p. 342, pl.
40, figs. 14-16 (not fig. 13), 1884.—Haruster, Abh. Schweiz. Pal. Ges., vol.
17, p. 64, pl. 10, figs. 9, 11, 1890—Eacrr, Abh. bay. Akad. Wiss. Miinchen,
Cl. II, vol. 18, p. 265, pl. 5, figs. 25, 26 (?), 1893——CHapMan, Proc. Zool.
Soe. London, 1895, p. 18—Goks, Bull. Mus. Comp. Zool., vol. 29, p. 33,
1896.—MIiLLert, Journ. Roy. Micr. Soc., 1899, p. 864.—Baae, U. S. Geol.
Surv. Bull. 514, p. 34, pl. 7, figs. 2a, b, 1912—Heron-ALLEN and HArLanp,
British Antarctic Exped., Zoology, vol. 6, 118, 1922.
Haplophragmoides trullissata, CUSHMAN, U. S. Nat. Mus. Bull. 71, pt. 1, p. 100,
figs. 148 a, 6 (in text), 1910.—Prarcry, Trans. Roy. Soc. Edinburgh, vol. 49,
p. 1008, 1914.—CusHMAN, U. S. Nat. Mus. Bull. 104, pt. 2, p. 43, pl. 9, fig. 5,
1920.
Test small, planispiral, composed of about three coils, not com-
pletely involute, the chambers of earlier coils visible at the center in
the umbilical region, which is also depressed, periphery slightly lob-
ulated; chambers numerous, 7 to 9 in the last-formed coil, subglobu-
lar; sutures distinct, slightly depressed ; wall of fine sand grains with
an excess of yellowish-brown or reddish-brown cement, smooth and
polished; aperture a short narrow slit slightly above the base of the
chamber; color yellowish brown or reddish brown. Diameter, 0.35
mm.; thickness, 0.1 mm.
There is a single very beautifully preserved specimen from A/da-
tross Station H3779, latitude 24° 45’ N., longitude 130° 16’ W., in
2,627 fathoms. This is the only record for the species or for the
family in the entire collection. The species is widely distributed in
the deep water of both the Atlantic and Pacific.
Family AMMODISCIDAE
Genus AMMODISCUS Reuss, 1861
AMMODISCUS INCERTUS (d’Orbigny)
There is a single very small specimen from Albatross Station
D3837, latitude 16° 32’ S., longitude 148° 40’ W., in 2,363 fathoms.
No other material was found in all the samples examined.
6 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Family TEXTULARIIDAE
Subfamily SPIROPLECTAMMININAE
Genus SPIROPLECTAMMINA Cushman, 1927
Spiroplectammina CusHMAN, Contr. Cushman Lab. Foram. Res., vol. 3, p. 23,
1927 ; Cushman Lab. Foram. Res. Spec. Publ. No. 1, p. 114, 1928.
Teztularia (part) PARKER and JONES (not Defrance), Philos. Trans., vol. 155,
p. 370, 1865.
Spiroplecta H. B. Brapy (not Ehrenberg), Rep. Voy. Challenger, Zoology, vol. 9,
p. 376, 1884.
Genoholotype.—Textularia agglutinans var. biformis Parker and
Jones.
Test free, early chambers planispiral in both microspheric and
megalospheric forms, later ones biserial; wall arenaceous with a yel-
lowish-brown cement; aperture in the planispiral portion at the base
of the apertural face, in the biserial portion at the inner margin of
the chamber. Carboniferous to Recent.
Out of all the Zewtularia group present in this collection from the
Pacific only mlletti seems to belong to this genus. At this time com-
ment may be made on the relationships of this genus to 7’extularia.
Doctor Lacroix has recently published a paper on the relationships
of these, seeming to show that “ Spiroplecta wrightii” is the same as
“Textularia sagittula.” We also comes to the conclusion that all
species of 7extularia have a spiral beginning, and that the chambers
may have been displaced where they are not present. An examina-
tion of many species of this genus does not carry out this hypothesis.
There are forms that have been placed under Spiroplectammina in
which both the megalospheric and microspheric forms have plani-
spirally coiled chambers in the end. These have been included under
Spiroplectammina. There are, however, a great many species excel-
lently preserved both in fossil and Recent material in which the
megalospheric form does not show any trace of spiral beginnings.
These seem to be distinct from the former group. It may be noted
here also that certain of the species referred to J'extularia evidently
have their relationship with Gaudryina, and the microspheric form
shows traces of a triserial beginning instead of a spiral one. From
a strictly phylogenetic point of view, these should be distinguished
from the other 7'ewtularia group having their beginning in a spiral
form. By acceleration of development the triserial stage has become
more and more restricted, and the biserial stage is taken on earlier
and earlier. This process may continue until the relationship with
Gaudryina is lost entirely, and there may be no possible way of dis-
tinguishing textularian forms developed from this source from those
that have developed from spiral beginnings.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS e
SPIROPLECTAMMINA MILLETTI (Cushman)
PLATE 1, Fiaures 5 a, b
Teatularia milletti CUSHMAN, U. S. Nat. Mus. Bull. 71, pt. 2, p. 183, figs. 18, 19
(in text), 1911.—HerRoN-ALLEN and Har.Anp, British Antarctic Exped.,
Zoology, vol. 6, p. 118, 1922.
Test compressed, somewhat longer than broad, composed of numer-
ous broad, low chambers, the whole test rapidly increasing in width,
especially in the earlier portion, end view narrow, periphery acute,
each chamber with the outer margin thickened and coarsely arena-
ceous, the early portion concave and much less roughened, so that
the surface of the test is composed of an alternating series of raised,
rough ridges, and smoother depressions; sutures rather indistinct
except for the depressions; wall coarsely arenaceous; aperture nar-
row at the inner margin of the last-formed chamber, with a slightly
overhanging lip. Length, 0.8 mm.
This species, which has been recorded from a number of stations in
the North Pacific and also by Heron-Allen and Earland from the
Antarctic, seems to have a wide distribution in the Pacific region.
It is nearest to a species described from the Atlantic as Zewxtularia
mexicana. 'The microspheric form shows distinct traces of coiling in
the early chambers.
The species has been recorded by Hofker from the Zuider Zee, but
from his figures his species from the Atlantic is not the same as this
one, which is well developed in the Pacific.
Subfamily TEXTULARIINAE
Genus TEXTULARIA Defrance, 1824
Tectularia DEFRANCE, Dict. Sci. Nat., vol. 82, p. 177, 1824—CusHMAN, Cushman
Lab. Foram. Res. Spec. Publ. No. 1, p. 144, 1928.
Textilaria BHRENBERG, Abh. kais. Akad. Wiss. Berlin, 1839, p. 135.
Piecanium ReEvss, Sitz. Akad. Wiss. Wien, vol. 44, p. 383, 1861 (1862) (genotype,
by designation, Textilaria labiata Reuss).
Grammostomum (part) of authors.
Palaeotextularia ScHuBERT, Pal. Zeitschr., vol. 3, p. 188, 1920 (genotype, by
designation, Textularia jonesi H. B. Brady).
Genoholotype—Textularia sagittula Defrance.
Test free, elongate, tapering, usually compressed with the zigzag
line between the chambers on the middle of the flattened sides, early
chambers in the microspheric form usually planispirally coiled, later
biserial, chambers simple, not labyrinthic; wall arenaceous, the rela-
tive quantity of cement varying much; aperture, typically an arched
slit at the inner margin of the chamber, occasionally in the apertural
face.
Cambrian to Recent.
8 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
TEXTULARIA FOLIACEA Heron-Allen and Earland
PLaTH 1, Figures 6-10
Textularia foliacea HERON-ALLEN and HARLAND, Trans. Zool. Soc. London, vol.
20, pt. 2, p. 628, pl. 47, figs. 17-20, 1915—CusHMan, U. S. Nat. Mus. Bull.
100, vol. 4, p. 117, pl. 19, figs. 7 a, b, 1921; Carnegie Inst. Washington Publ.
342, p. 14, pl. 2, figs. 24; pl. 3, fig. 1, 1924.
Test of medium size, much compressed, triangular, the outer aper-
tural end broadly convex; chambers numerous, 9 to 11 on each side
in the adult; sutures fairly distinct but only slightly depressed;
surface roughly formed, of sand grains so cemented as to leave a
characteristic, rough, somewhat flaky surface; aperture fairly large,
curved, at the base of the last-formed chamber, with a slight lip.
Length, 1 to 1.5 mm.
This species is one of the commonest in the shallow water of the
general Indo-Pacific region. It is much compressed and has a char-
acteristic size and shape in the adult specimens. When seen with
other specimens in the sample it is usually much lighter in color.
It is evidently most at home in shallow water of coral-reef regions,
and while there are some records from deeper water they are mostly
small specimens that may have been easily carried into deeper water.
Previous records are by Heron-Allen and Farland from the Kerimba
Archipelago, from Timor Sea, Java, 50 fathoms. I have recorded it
from seven stations off the Philippines at depths ranging from 20 to
494 fathoms, and from shallow water about Samoa. In the present
collections it has occurred at the following localities: Mokaujar
Anchorage, Fiji; Viva Anchorage, Fiji, 3 fathoms; Levuka, Fiji, 12
fathoms; Nairai, Fiji, 12 fathoms and 24 fathoms; Vavau Anchorage,
Tonga Islands; Rongelap Atoll, Marshall Islands; and from the
Guam Anchorage, Ladrone (Marianas) Islands, 21 fathoms. At the
last locality specimens are abundant, and the largest in size of any
of the localities.
TEXTULARIA FOLIACEA Heron-Allen and Earland OCEANICA, new variety
PLaTe 1, Ficures 11, 12
Variety differing from the typical in the much thicker form and
rougher appearance of the test. Length of holotype of variety, 2
mm.; width, 1.25 mm.; thickness, 1 mm.
Holotype of variety—Cushman Coll. No. 14667, from Mokaujar
Anchorage, Fiji.
This variety is much thicker than the typical, much rougher, and
usually of a much darker color. The two occur together at some of
the localities, but the variety seems to occur alone at certain others.
The holotype is an exceptionally large, fine specimen, much larger
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 9
than the usual run of specimens in the collection. In the early stages
the variety is usually distinguished by its greater thickness.
Besides the type locality the variety occurs at Levuka, Fiji, 12
fathoms; Nairai, Fiji, 12 fathoms; Vavau Anchorage, Tonga Islands;
Rotonga, 7 fathoms; and Port Lotten, Kersail, Caroline Islands.
TEXTULARIA SEMIALATA Cushman
PLATE 2, Ficurns 1-3
Texrtularia semialata CUSHMAN, Proc. U. S. Nat. Mus., vol. 44, p. 634, pl. 80,
figs. 6, 7, 1913; U. S. Nat. Mus. Bull. 100, vol. 4, p. 116, pl. 24, figs. 2, 3,
1921.
Test compressed, rather rapidly increasing in width, the periphery
rounded, especially in the later chambers; chambers numerous, con-
siderably overlapping so that all except the last two are much
broader than high from the exterior; sutures distinct, nearly hori-
zontal, depressed; wall finely arenaceous, very smoothly finished;
aperture at the inner margin of the chamber with a distinct, over-
hanging lip. Length, 0.8 mm.; breadth, 0.55-0.7 mm.; thickness,
0.8-0.35 mm.
This species was originally described from the Philippines. Very
typical specimens of the broad form occur at Mokaujar Anchorage,
Fiji, and a single specimen, apparently the same, from Albatross
Station H3901, in 1,620 fathoms, off Marokau Island, Paumotus
(Tuamotu Archipelago).
TEXTULARIA CANDEIANA d’Orbigny
PLATE 2, Fiaures 4 a, 0b
Textularia candeiana v’ORBIGNY, in De la Sagra, Hist. Fis. Pol. Nat. Cuba,
“Foraminiféres,” p. 148, pl. 1, figs. 25-27, 1839.—Fornasini, Mem. Accad.
Sci. Istit. Bologna, ser. 5, vol. 10, p. 303, pl. 0, fig. 8, 1908.—CHAPMaN, Rep.
Foram. Subantarctic Islands, New Zealand, p. 329, 1909—CusuMan, U. S.
Nat. Mus., Bull. 71, pt. 2, p. 12, figs. 14-17 (in text), 1911.—Hrron-ALLEN
and EHarLANpD, Trans. Zool. Soc. London, vol. 20, pt. 2, p. 627, pl. 47, figs.
10-16, 1915; Trans. Linn. Soc. London, vol. 11, ser. 2, p. 230, pl. 41, figs. 1, 2,
1916; Journ. Roy. Micr. Soc., 1916, p. 41.—CusuHman, Carnegie Inst.
Washington Publ. 291, p. 32, 1922; Proc. U. S. Nat. Mus., vol. 59, p. 50,
pl. 11, figs. 7, 8, 1921; U. S. Nat. Mus. Bull. 100, vol. 4, p. 109, 1921; Carne-
gie Inst. Washington Publ. 311, p. 32, pl. 2, fig. 2, 1922: U. S. Nat. Mus.
Bull. 104, pt. 3, p. 8, pl. 1, figs. 1-8, 1922—Hrron-Atten and EaRLanp,
British Antarctic Expedition, Zoology, vol. 6, p. 119, 1922; Journ. Linn.
Soc. Zool., vol. 35, p. 618, 1924—HaAnzawa, Jap. Journ. Geol. Pal., vol. 4,
p. 88 (table), 1925 (1926).—CusHMAN, Carnegie Inst. Washington Publ.
344, p. 76, 1926.
Textularia sagittula DE¥YRANCE yar. candeiana Mittert, Journ. Roy. Micr. Soc.,
USO) Ps oboe, pl. ts fiz. 2.
10 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Test elongate, club-shaped, the early portion narrow, much com-
pressed, the edges almost carinate, tapering gradually to the rounded
apex; chambers numerous, those of the early portion somewhat com-
pressed, later ones enlarging very rapidly, and the final ones becom-
ing much inflated; sutures of the later portion fairly distinct and
considerably oblique, very slightly depressed; wall rather coarsely
arenaceous especially in the early portion, often obscuring the
sutures; aperture in a broad, shallow sinus at the base of the inner
margin of the chamber. Length, up to 1 mm.; breadth, up to 0.6
mm.; thickness, 0.5 mm.
This species is common in the West Indian region, from which it
was originally described by d’Orbigny. It occurs also in typical
form in the Pacific. It may be distinguished from the following
species (agglutinans) by the greater obliquity of the sutures and by
the very rapid expansion of the chambers toward the apertural end
together with the characteristic roughness of the surface of the early
chambers. In the Pacific material examined this species and 7’.
agglutinans occur together very rarely, the latter occurring in the
Fiji collections but the former apparently absent.
Our records for 7. candeiana include the following localities:
Makemo Lagoon, Paumotus; Rangiroa; Rutavu; Port Lotten, Ker-
sail, Caroline Islands; Rongelap Atoll, Marshall Islands; Guam
Anchorage, Ladrone Islands, 21 fathoms; and the following Alba-
tross stations: H8814, latitude 15° 14’ 10’ S., longitude 147° 51’
05’ W., 391 fathoms off the Paumotus; H3855, off Apataki Island,
Paumotus, 654 fathoms; H38889, off Taenge Atoll, Paumotus, 928
fathoms; and H3898, off Hikueru Atoll, Paumotus, 348 fathoms.
TEXTULARIA AGGLUTINANS d’Orbigny
PLATE 2, FicuRES 5-7
Textularia agglutinans v’ORBIGNY, in De la Sagra, Hist. Fis. Pol. Nat. Cuba,
“ Foraminiféres,” p. 136, pl. 1, figs. 17, 18, 32, 34, 1839.—H. B. Brapy, Rep.
Voy. Challenger, Zoology, vol. 9, pl. 48, figs. 1 a, b (var. 2, 3), 1884.—Goks,
Bull. Mus. Comp. Zool., vol. 29, p. 41, 1896.—FLinr (part), Rep. U. S. Nat.
Mus., 1897, p. 284, pl. 29, fig. 4, 1899CusHmaAn, U. S. Nat. Mus. Bull. 71,
pt. 2, p. 9, figs. 10 a, b (in text), 1911; Proc. U. S. Nat. Mus., vol. 59, p. 49,
pl. 11, figs. 1-8, 1921; U. S. Nat. Mus. Bull. 100, vol. 4, p. 106, pl. 20, fig. 8,
1921; Carnegie Inst. Washington Publ. 311, p. 22, pl. J, fig. 6, 1922; U. S.
Nat. Mus. Bull. 104, pt. 3, p. 7, pl. 1, figs. 4, 5, 1922; Carnegie Inst. Wash-
ington Publ. 344, p. 76, 1926.
Test elongate, tapering, very little compressed, periphery rounded,
the increase in diameter from the initial end rather uniform; cham-
bers inflated, increasing somewhat in height toward the apertural
end rather uniformly; sutures distinct, depressed, usually about at
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 11
right angles to the long axis of the test; wall rather coarsely arena-
ceous, but smoothly finished; aperture an elongate slit in a well-
marked depression of the inner border of the chamber. Length,
0.8-1 mm.; breadth, 0.5-0.6 mm.
This species was described by d’Orbigny from shore sands of Cuba,
and it is a typical species of the general West Indian region, occur-
ring in typical form at least as far back as the Miocene in Florida
and probably in the West Indies. It is not so common in the Pacific
but occurs widely distributed, especially in rather shallow water.
It may be distinguished from candeiana by the horizontal sutures,
the uniform tapering test, and the rather uniform shape of the cham-
bers. In the Pacific collections it has occurred at Albatross Station
H3961, off the south coast of Tarawa, Gilbert Islands, 4138 fathoms.
In the shallow water material it occurs off the Fiji Islands from
Levuka, 12 fathoms, Mokaujar Anchorage. There are typical speci-
mens also from Rongelap Atoll, Marshall Islands. It is much less
common than candeiana.
TEXTULARIA CONICA d’Orbigny
PLATE 2, FIGURES 8-10; PLATE 3, FIGURES 1, 3
Textularia conica D’ORBIGNY, in De la Sagra, Hist. Fis. Pol. Nat. Cuba, “ Foram-
iniféres,” p. 143, pl. 1, figs. 19, 20, 1839.—H. B. Brapy, Rep. Voy. Chal-
lenger, Zoology, vol. 9, p. 365, pl. 48, figs. 138, 14; pl. 118, fig. 1, 1884.—
Goks, Bull. Mus. Comp. Zool., vol. 29, p. 48, 1896—CusHMAN, Proc. U. 8S.
Nat. Mus., vol. 59, p. 50, pl. 11, figs. 46, 1921; U. S. Nat. Mus. Bull. 100,
vol. 4, p. 128, pl. 25, figs. 2 a—c, 1921; Carnegie Inst. Washington Publ. 311,
p. 24, pl. 2, fig. 4, 1922; U. S. Nat. Mus. Bull. 104, pt. 3, p. 22, pl. 5, figs. 5-7,
1922; Carnegie Inst. Washington Publ. 342, p. 15, pl. 1, fig. 6, 1924.
Test usually wider than high, triangular in front view, broadly
oval in end view, slightly compressed, apex bluntly pointed; cham-
bers comparatively few, distinct; sutures distinct, slightly depressed ;
wall arenaceous, smooth, or slightly roughened; aperture a narrow
slit at the base of the inner margin of the last-formed chamber;
color gray. Length, 0.55-0.7 mm.; breadth, 0.55-0.65 mm. ; thickness,
0.4-0.5 mm.
D’Orbigny’s original types of this species were from the West
Indies in shallow water, where it is common and widely distributed.
It has been recorded from many regions both as a Recent and fossil
species, but an examination of the records where figures are given
would show that these are not all one species. Various authors have
referred the species to Textularia abbreviata d’Orbigny described by
him from the Vienna Basin Miocene, but that species in its typical
form is very different from the Recent West Indian one. The species
occurs in our material from Rangiroa, Paumotus; Vavau Anchorage,
91518—32-——_2
12 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Tonga Islands; Mokaujar Anchorage, Fiji Islands; and from Ron-
gelap Atoll, Marshall Islands. There is also a typical specimen from
Albatross Station H8930, off Anu Anuraro Atoll, Paumotus, 438
fathoms, evidently a specimen carried out into deep water, as the
station is only three-fourths of a mile offshore.
TEXTULARIA CORRUGATA Heron-Allen and Earland
PLATE 3, FIGURES 2, 4
Textularia conica pD’ORBIGNY var. corrugata HERON-ALLEN and HARLAND, Trans,
Zool. Soe. London, vol. 20, p. 629, pl. 47, figs. 24-27, 1915.
Test short and broad, nearly as broad as long, oval in end view;
chambers very distinct, much excavated toward the base, low and
broad; sutures deep, usually curved upward in the middle, thence
rather sharply downward toward the periphery; wall arenaceous,
fine, of rounded particles with considerable cement giving it a smooth
finish. Length and breadth, up to 1 mm.; thickness, 0.65-0.75 mm.
Heron-Allen and Earland described this species from Kerimba
Archipelago off southeast Africa as a variety of Teatularia conica.
While the two forms occur together occasionally in our material they
seem to be very distinct. This species is widely distributed, occur-
ring off the Fiji Islands in 40 to 50 fathoms, off Rongelap Atoll,
Marshall Islands, and Guam Anchorage, Ladrone Islands, in 21
fathoms.
TEXTULARIA ALBATROSSI Cushman
PLATE 3, Ficurnes 5 a, 0
Textularia concava FLinT (part) (not Karrer), Rep. U. S. Nat. Mus., 1897,
p. 283, 1899.
Texrtularia albatrossi CusHMAN, U. S. Nat. Mus. Bull. 104, pt. 3, p. 14, pl. 2,
figs. 5, 6, 1922.
Test elongate, much tapering, apical end bluntly pointed, apertural
end broadly angled, in the later portion the breadth nearly equal
to the thickness, concave at each side in the middle, periphery con-
vex; chambers distinct, the last ones much inflated, low and broad,
each with a reentrant portion near the central part at each side;
sutures distinct, depressed; wall coarsely arenaceous, rather smoothly
finished, especially in the later portion; aperture a long, narrow slit
above the base of the chamber, with a slight lip; color gray. Length,
0.9 mm.; breadth, 0.55 mm.; thickness, 0.4 mm.
Previous records of this species are from the Atlantic, where it
occurs at a depth of 382 fathoms in the western part of the Carib-
bean Sea. The single specimen figured here is the only one that
occurs in our material, and is from Albatross Station H8858, Ngaruae
Pass, Fakarava Atoll, Paumotus, in 599 fathoms.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 13
TEXTULARIA CONCAVA Karrer
PLATE 3, WiaeuREes 6 a, 0
Many forms have been referred to this species described by Karrer
from the Miocene of Europe. Some of the specimens figured from
the Mediterranean are very close to Karrer’s original figure, and
also some Recent Indo-Pacific shallow water specimens. The only
specimen in the present collection that can be referred to Karrer’s
species is that figured, which is from Albatross Station H3815,
latitude 15° 15’ 15’’ S., longitude 147° 51’ 35” W., in 524 fathoms
Family VERNEUILINIDAE
Genus GAUDRYINA d’Orbigny, 1839
Gaudryina D’OrBIGNy, in De la Sagra, Hist. Fis. Pol. Nat. Cuba, “ Foraminif-
éres,” p. 109, 1889.—CusHmMaAN, Cushman Lab. Foram. Res. Spee. Publ.
No. 1, p. 126, 1928.
Genotype—By designation, Gaudryina rugosa d’Orbigny.
Test in the early stages triserial, later biserial, transverse section
angled or rounded; wall coarsely or finely arenaceous often with a
large proportion of cement; aperture, typically a low opening at
the base of the inner margin of the chamber, or in the wall of the
apertural face. Lower Cretaceous to Recent.
There are a number of species in this Pacific material, and these
show very little difference in the relative amount of the test made
up by the triserial portion. As already noted under Spiroplectam-
mina, textularian forms may easily be developed from Gaudryina
and the triserial portion entirely crowded out. All stages in this
process can be seen in various species of the genus.
GAUDRYINA BRADYI Cushman
PLATE 3, Ficures 8, 9
Gaudryina pupoides H. B. Brapy (not G. pupoides d’Orbigny), Rep. Voy.
Challenger, Zoology, vol. 9, p. 878, pl. 46, figs. 14, 1884——H. B. Brapy,
PaRKER, and JONES, Trans. Zool. Soc. London, vol. 12, p. 219, pl. 43, figs. 7,
8, 1888.—Wericut, Ann. Mag. Nat. Hist., ser. 6, vol. 4, p. 448, 1889.—
Prarcry, Trans. Nat. Hist. Soc. Glasgow, vol. 2, p. 176, 1890.—WricuHt,
Proc. Roy. Irish Acad., ser. 3, vol. 1, p. 471, 1891.—CHapMan, Proc. Zool.
Soe. London, 1895, p. 20.—Goiis, Bull. Mus. Comp. Zool., vol. 29, p. 40,
1896.—F.LInT, Rep. U. S. Nat. Mus., 1897, p. 287, pl. 32, fig. 4, 1899.—
CHAPMAN, Journ. Linn. Soe. Zool., vol. 30, p. 403, 1910; Zool. Res. Hn
deavour, pt. 3, p. 310, 1912; vol. 3, pt. 1, p. 16, 1915.—Siprsortom, Journ.
Roy. Micr. Soc., p. 28, 1918.
Gaudryina bradyi CusuMan, U. S. Nat. Mus. Bull. 71, pt. 2, p. 67, figs.
107 a-c (in text), 1911.—Prarcey, Trans. Roy. Soc. Edinburgh, vol. 49, p.
1014, 1914—CusuMaAn, U. S. Nat. Mus. Bull. 100, vol. 4, p. 149, pl. 29, fig. 3,
1921; U. S. Nat. Mus. Bull. 104, pt. 3, p. 74, pl. 12, fig. 8, 1922.
14 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Test stout, somewhat elongate, tapering slightly until near the
initial end, where it tapers abruptly to the somewhat blunt end;
triserial portion nearly circular in cross section, of few chambers,
the later biserial portion making up about three-fourths of the test,
slightly compressed; chambers overlapping and appearing crowded,
broadly elliptical in cross section, inflated; sutures deep and distant,
end strongly convex; wall of fine arenaceous or calcareous shell ma-
terial, smooth; aperture oval, slightly back from the inner margin
of the chamber and with the border raised somewhat and thick-
ened; color light gray. Length, 0.88-1 mm.
Although this is a very widely distributed species, it is very rare
in this collection. Single specimens, both of which are here figured,
occurred at Albatross Stations H3818, latitude 15° 24’ 10” S., longi-
tude 147° 56’ W., in 897 fathoms, and H38901, 8 miles off Marokau
Islands, Paumotus, 1,620 fathoms.
GAUDRYINA TRIANGULARIS Cushman var. ANGULATA Cushman
PLATE 3, FIGURES 7 @, 0
Gaudryina triangularis CUSHMAN var. angulata CUSHMAN, Carnegie Inst. Wash-
ington Publ. 342, p. 22, 1924.
Variety differing from the typical in the later chambers, which are
very angular, the opposite sides parallel, giving a peculiar appear-
ance to the test, especially in end view. Length, 0.6 mm; breadth,
0.45 mm; thickness, 0.38 mm.
This variety, originally described from Samoa, occurs rarely in
the Fiji Islands at Mokaujar Anchorage.
GAUDRYINA QUADRANGULARIS Bagg
PLATE 38, FIGURES 10, 11
Gaudryina quadrangularis Bacc, Proc. U. S. Nat. Mus., vol. 34, p. 133, pl. 5,
fig. 1, 1908 —CusuMan, U. S. Nat. Mus. Bull. 71, pt. 2, p. 64, fig. 103 (in
text), 1911; Proc. U. S. Nat. Mus., vol. 56, p. 604, 1919; U. S. Nat. Mus.
Bull. 100, vol. 4, p. 147, pl. 29, fig. 2, 1921.
Test elongate, tapering abruptly at the initial end, composed of an
early trihedral portion with acute angles made up of a triserial group
of chambers, and a later more or less quadrangular portion composed
of chambers biserially arranged; sutures often largely obscured by
roughening of the surface; wall coarsely arenaceous and rather
roughly cemented; aperture an elongate orifice between the inner
border of the chamber and the adjacent wall of the preceding cham-
ber, with, in end view, a sort of rounded lip above and a depression
at either side; in side view the aperture is in a depression of the
abruptly truncated end of the test. Length, up to 3 mm.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 15
This species was originally described by Bagg from off the
Hawaiian Islands. It is now known from numerous stations, espe-
cially from the Philippine region and in the North Atlantic. It has
proved to be the most common species of the genus in these collec-
tions but occurs only in the Albatross material from comparatively
deep water as follows: Station H3808, entrance to Avatoru Pass,
Rahiroa Atoll, Paumotus, 645 fathoms; H3810, same locality, 661
fathoms; H3959, Ngaruae Pass, Fakarava Atoll, 666 fathoms; H3860,
off Fakarava, 602 fathoms; and H38954, off Nomuka Island, Cook
Islands, 600 fathoms.
The specimens are uniform in showing the peculiar flaky rough-
ness, especially over the earlier chambers, obscuring the sutures to a
considerable degree.
GAUDRYINA sp. (7)
PLATE 4, FIGURES 2 a, Db
The peculiar form figured here occurred in Makemo Lagoon,
Paumotus. No other specimens were found with this to give the
complete characters of the species.
GAUDRYINA RUGULOSA, new species
PLATE 4, F1cuRES 1 a, D
Test large, stout, somewhat longer than broad, early triserial
portion usually confined to the earliest stages, later becoming biserial
for most of the growth, periphery broadly rounded; chambers fairly
distinct, with the lower side much excavated, sometimes with one
or more backwardly directed projections overlapping onto the pre-
ceding chamber; sutures distinct, depressed, nearly horizontal or
somewhat upwardly curved in the middle; wall arenaceous, but with
much cement rather smoothly finished; aperture a low opening at
the inner margin of the last-formed chamber. Length, 2 mm.;
breadth, 1.85 mm.; thickness, 0.75 mm.
Holotype.—Cushman Coll. No. 14698, from Alofi Niue.
This species has been often recorded from the Indo-Pacific as
Textularia rugosa (Reuss). The early stages are, however,
triserial, and the species should be placed in Gaudryina. This has
been noted by Heron-Allen and Earland in their Kerimba paper,
and shown by them in figures in that same paper (pl. 47, figs. 7-9).*
Though the species described by Reuss from the Tertiary from
Gaas has apparently chambers that are excavated somewhat at the
base, it is certainly not the same as this very large species, so
1 The Foraminifera of the Kerimba Archipelago, Trans. Zool. Soc. London, vol. 20, pt. 2,
pp. 543-794, pls. 40-53, 1915.
16 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
well developed in the Indo-Pacific in shallow water of coral reefs.
Besides the type locality where the species is common and well de-
veloped, it has also occurred in the material from Guam Anchorage,
Ladrone Islands, 21 fathoms. It occurs in the Philippines, where
it is often common and grows to large size.
Genus CLAVULINA d’Orbigny, 1826
Clavulina p’OrBiaeny, Ann. Sci. Nat., vol. 7, p. 268, 1826—CusHMan, U. S. Nat.
Mus. Bull. 104, pt. 3, p. 80, 1922; Cushman Lab. Foram. Res. Spec. Publ.
No. 1, p. 127, 1928.
Verneuilina (part) of authors.
Valvulina (part) of authors.
Tritaxia (part) of authors.
Genotype.—By designation, Clavulina parisiensis d’Orbigny.
Test elongate, cylindrical or angled, early portion consisting of
chambers arranged triserially, in most species quickly followed by
a uniserial series, but in some primitive species with a biserial stage
between; wall arenaceous; aperture, in the early stages, a simple
opening at or near the inner margin, later becoming terminal, often
with a neck, in some species with an apertural tooth.
Cretaceous to Recent.
CLAVULINA CGMMUNIS d’Orbigny
PLATE 4, Figures 3 a, 0
There is a single large specimen figured here that may be referred
to this species. It is from Albatross Station H3816, latitude 15° 16’
50’’ S., longitude 147° 52’ 30’ W., in 450 fathoms. The typical
form of the species is often common in shallow water of the Mediter-
ranean, and it is surprising that it has not occurred in many of the
shallow-water Pacific samples included in this paper.
CLAVULINA PACIFICA Cushman
Puatp 4, Ficures 4, 7, 9
Clavulina angularis H. B. Brapy (in part) (not d’Orbigny), Rep. Voy. Chailen-
ger, Zoology, vol. 9, p. 396, pl. 48, figs. 22-24, 1884.
Clavulina pacifica CUSHMAN, Carnegie Inst. Washington Publ. 342, p. 22, pl.
6, figs. 7-11, 1924.
Test elongate, early portion triserial, short, trihedral; later por-
tion uniserial, the sides parallel or nearly so, triangular in transverse
section, the sides flat or somewhat convex; chambers fairly high;
sutures distinct, depressed, curved; wall arenaceous, but very
smoothly finished, last-formed chamber tapering toward the aper-
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 17
tural end, the whole rounded; aperture circular, with a fairly large
tooth; color yellowish brown. Length, up to 1.5 mm.
This species was described from Samoa, where it occurred at sev-
eral stations. The species has been usually recorded from the Pacific
as Clavulina angularis. It is, however, a very different species from
that of d’Orbigny and also from C. tricarinata, which is characteristic
of the West Indian region. It is apparently not so common in the
shallow water of the Pacific as the following species, C. difformis H.
B. Brady. Typical specimens of (. pacifica were obtained from
Rangiroa and from Makemo Lagoon, Paumotu Islands, specimens
from both localities being figured here.
CLAVULINA DIFFORMIS H. B. Brady
~
PLatTE 4, Ficures 5, 6, 8, 10
Clavulina angularis D’ORBIGNY var. difformis H. B. Brapy, Rep. Voy. Challenger,
Zoology, vol. 9, p. 896, pl. 48, figs. 25-81, 1884—Hrron-ALLEN and HARLAND,
Trans. Zool. Soc. London, vol. 20, pt. 2, p. 637, pl. 48, figs. 20-22, 1915.—
CusHMAN, U. S. Nat. Mus. Bull. 100, vol. 4, p. 156, pl. 31, figs. 2 a, b, 1921.
Clavulina difformis CUSHMAN, Carnegie Inst. Washington Publ. 342, p. 23,
pl. 6, figs. 5, 6, 1924.
Test elongate, somewhat tapering, usually polygonal, the sides con-
cave, early triserial portion short, angles of the test sharp, slightly
produced; chambers numerous, distinct, slightly inflated; sutures
distinct, depressed, curved; wall arenaceous, rather roughly finished ;
aperture circular, with a distinct tooth, the last-formed chamber
short and at the apertural end broadly truncate or even slightly de-
pressed. Length, up to slightly more than 1 mm.
Brady described this as a variety of Clavulina angularis. It is,
however, a distinct species developed in the Indo-Pacific region.
The test is typically polygonal in end view, and the wall is very dis-
tinctly roughened, the chambers slightly inflated and often having
distinct angular projections, which overlap the preceding chamber.
Previous records include Nares Harbor, Admiralty Islands (Brady) ;
Kerimba Archipelago, southeast coast of Africa (Heron-Allen and
Farland) ; Apra Bay, Guam, and Samoa (Cushman). In the present
collections typical specimens occurred at Guam Anchorage, Ladrone
Islands, 21 fathoms; Mokaujar Anchorage, Fiji Islands, Vavau An-
chorage, Tonga Islands; and Makemo Lagoon, Paumotu Archipelago.
Family MILIOLIDAE
Genus QUINQUELOCULINA d’Orbigny, 1826
Quinqueloculina D’OrBieny, Ann. Sci. Nat., vol. 7, p. 301, 1826—CusHMAN, U. S.
Nat. Mus. Bull. 71, pt. 6, p. 42, 1917; Cushman Lab. Foram. Res. Spec. Publ.
No. 1, p. 146, 1928; U. S. Nat. Mus. Bull. 104, pt. 6, p. 21, 1929.
18 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Serpula (part) LinNAEus, Systema naturae, ed. 10, p. 786, 1758.
Adelosina p’OrBIGNyY, Ann. Sci. Nat., vol. 7, p. 803, 1826 (genotype, by designa-
tion, A. striata d’Orbigny).
Uniloculina D’OrBIGNY, Foram. Foss. Bass. Tert. Vienne, p. 261, 1846 (genoholo-
type, U. indica d’Orbigny).
Miliolina (part) Wut~rAmMson, Rec. Foram. Great Britain, p. 83, 1858 (and
later authors).
Genotype.—By designation, Serpula seminulum Linnaeus.
Test with the coiling in five planes, the chambers half a coil in
length and added successfully in planes 144° apart, five chambers
completing a cycle, each chamber 72° from its adjacent one but 144°
from the immediately preceding one; wall imperforate, calcareous,
often with arenaceous material on the exterior and in deep or brack-
ish water occasionally becoming siliceous; aperture usually with a
simple tooth.
This is one of the common genera in the shallow water of the South
Pacific represented by numerous species. In addition to those recorded
here, there are probably several others in our material, but they are
represented by either incomplete or insufficient specimens so that
specific determination of them is not possible. Some of the species
show considerable variation in shape of the chambers and the orna-
mentation from the young stages to the adult, and it is necessary to
have rather full suites of specimens in order really to understand
the complete developmental stages and the adult as well. In order to
facilitate use of the species of the genus they are here arranged
alphabetically.
QUINQUELOCULINA ANGUINA Terquem var. AGGLUTINANS (Wiesner)
'
PLatTH 5, Ficures 1 ac
Miliolina anguina (TERQUEM) var. dgglutinans WIESNER, in Heron-Allen and Har-
land, Trans. Zool. Soc. London, vol. 20, pt. 2, p. 575, 1915—CusHMAN, Car-
negie Inst. Washington Publ. 342, p. 60, pl. 22, figs. 5, 6, 1924.
Test elongate, two or three times as long as broad, periphery
rounded; chambers distinct, rounded in transverse section, the aper-
tural end extended; sutures somewhat depressed; wall with the sur-
face coarsely arenaceous and roughly finished; aperture with a
cylindrical neck, and slight lip. Length, 0.5 mm.; breadth, 0.3 mm.;
thickness, 0.2 mm.
This has proved to be a common form, well developed in our
material. Previous records for the Pacific are from the Kerimba
Archipelago off southeast Africa (Heron-Allen and Earland) and
from Samoa (Cushman).
Specimens occurred in considerable numbers at the following lo-
calities: Mokaujar Anchorage, Fiji Islands; Levuka, Fiji, 12
fathoms; off Nairai, Fiji, 12 fathoms; Vavau Anchorage, Tonga
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS
Islands, 18 fathoms; Rangiroa; Rotonga, 7 fathoms; Rongelap
Atoll, Marshall Islands; and Guam Anchorage, Ladrones, 21 fathoms.
QUINQUELOCULINA BERTHELOTIANA d’Orbigny
PLATE 5, Figures 2-4
Quinqueloculina berthelotiana p’ OrBiany, in Barker, Webb, and Berthelot, Hist.
Nat. Iles Canaries, vol. 2, pt. 2, “ Foraminiféres,” p. 142, pl. 3, figs. 25-27,
1839.
Test somewhat longer than broad, somewhat compressed; cham-
bers distinct, polygonal in section, the periphery concave, and the
TABLE 3.—Quinqueloculina berthelotiana—material examined
Num-
OSM Es os
mens
2107 (ooo 1 | H3309
21978__...- 4 | H3815
21979__--.- 3 | H3827
21980-_-_--- 1 | H3s34
219815 22-82 2 | H3848
21982__-__- 5 | H8858
21983 _ -___- 1 | H33859
21984. =~ 3 | H8875
21985___-_- 1 | H3876
21986. _ _--- 1 | H3884
21987. _---- 3 | H8891
21988____- - 8 | H3808
21989... -- 1 | H3889
21990_ = =. 1 | H8905
2190 Ieee ae 8 | H3910
21992_____- 2 | H3914
21993--...-.- 1 | H28916
21994. _____ 2 | H3926
21995____- = 3 | H3928
21996 _____- 4 | H3930
21997_____- 1 | H3931
21998_____- 6 | 3935
Locality
Entrance to Avatoru’ Pass,
Rahiroa Atoll, 2.5 miles 8.
Lat. 15° 15’ S., long. 147° 51’ 35”
W.
Lat. 14° 53’ 20’ S., long. 148° 42’
30” W.
West coast of Makatea Island,
1.3 miles E.
Village on west side Niau Atoll,
34 mile E.
Ngaruae Pass, Fakarava Atoll,
28° S., 1 mile E.
Ngaruae Pass, Fakarava Atoll,
35° S., 3.5 miles E.
Southwest point Tahanae, about
¥% mile offshore, 38 miles NE.
Northwest entrance Makemo
Lagoon, 1 mile SE.
Northwest point Raroia, 14 mile
SE.
Lat. 16° 30’ S., long. 148° 41’ W_-
Northwest point Hikueru Atoll,
44 mile E.
Northwest point Hikueru Atoll,
1.3 miles E.
Northwest point Hao Atoll, 4%
mile SE.
Southwest point Aki Aki, 1 mile
E.
Northeast point Nukutavake, 1
mile S.
Pinaki Atoll, 1 mile E_..--------
Midway between Nukutipipi
and Anu Anurunga.
Anu Anurunga, 1 mile SE_-___--
Anu Anuraro Atoll, $4 mile NW-
Anu Anuraro Atoll, 44 mile SE_
Hereheretue Atoll, 1 mile W_--_-
1 Key to abbreviations is given in Table 1.
Bot-
tom
tem-
pera-
ture
Depth
Faihoms| ° F.
645
508 | 40.2
540
348
39.7
43.8
798 | 37.8
425
377 | 43
636 | 38.9
486 | 41
1,609 | 35.5
659 | 38.5
438
405
40.7
42.5
594 | 39.5
Character of Occur-
bottom 1 rence
fne. wh, co. s___-| Rare.
wh. co. s. brk. sh.| Few.
ers. wh. co. s. | Rare.
vol. part.
wh. co. s, Mang. Do.
nod.
co. s. glob. oz....| Do.
Crs) CON S222 es Few.
pter. oz. vol. | Rare.
part.
CES: CO.#S- tees = Do.
whi co;si242222 Do.
ers. co. S. pter. Do.
OZ.
co. s. pter. oz..--| Do.
co. s. brk. sh--__| C 0 m -
mon.
co. s. pter. oz____| Rare.
ers! coy si 2S4.g- J! Do.
CONSE ores eS Com-
mon.
Copisee=ee Se- 22583 Rare
crs. co. s. pter. Do,
oz.
co.s. mang. glob_| Do.
co. s. brk. sh. Do.
pter. oz.
CO) Sr ee Few.
co. s. pter. oz. | Rare.
mang. part.
CTS: CONS. cease Few.
19
20 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
somewhat bluntly carinate edges of the test often becoming sinuous;
sutures fairly distinct, only slightly if at all depressed; wall smooth
or slightly roughened, with minute pits; aperture generally rounded
at the end of a distinct neck, and with a single short tooth. Length,
0.6-0.75 mm.; breadth, 0.3-0.4 mm.; thickness, 0.15-0.2 mm.
Many different forms similar to those shown in our figures have
been assigned numerous specific names. d’Orbigny’s originals seem
to be of the form that has polygonal chambers and a distinctly ex-
serted neck. The degree of sinuosity of the edges of the chambers
is very variable. Our specimens cover a wide range of territory
including a number of Albatross stations, data for which are given
in the accompanying table. From our other stations it occurs at
Rangiroa; Rutavu; 40 to 50 fathoms off Fiji Islands; Makemo
Lagoon; Hereheretue; Pinaki Atoll; Rongelap Atoll, Marshall
Islands; Port Lotten, Kersail, Caroline Islands.
QUINQUELOCULINA BIDENTATA d’Orbigny
PLATE 5, FIGURE 5
Quinqueloculina bidentata D’ORBIGNY, in De la Sagra, Hist. Fis. Pol. Nat. Cuba,
“ Foraminiféres,” p. 197, pl. 12, figs. 18-20, 1889.—CusHman, Proc. U. S.
Nat. Mus., vol. 59, p. 65, pl. 15, figs. 11, 12, 1921; Carnegie Inst. Wash-
ington Publ. 311, p. 54, 1922; Publ. 342, p. 59, pl. 22, figs. 1, 2, 1924.
Test somewhat longer than broad, periphery squarely truncate or
somewhat concave; chambers fairly distinct, rounded or polygonal
in transverse section; sutures somewhat obscure; wall composed on
the exterior of rather coarse sand grains, but fairly smoothly fin-
ished ; aperture somewhat extended with a slight lip and thin plate-
like tooth, which extends out beyond the opening. Length, 1 mm.;
breadth, 0.6 mm.
This species seems to be identical with that described by d’Orbigny
from the West Indian region. I have already recorded it from
Samoa. It is, however, not nearly so common in the region as are
the other two arenaceous forms. Specimens occurred only off the Fiji
Islands; Mokaujar Anchorage, Levuka, 12 fathoms; and Nairai
Islands, 12 fathoms.
QUINQUELOCULINA COSTATA d’Orbigny
PLATE 5, FIguRES 6, 7
Quinqueloculina costata p’OrBIaNy, Ann. Sci. Nat., vol. 7, p. 301, No. 3, 1826.—
TERQUEM, Mém. Soc. Géol. France, ser. 3, vol. 1, p. 68, pl. 6 (11), figs.
3 a-5 c, 1878.—CusHMAN, U. S. Nat. Mus. Bull. 71, pt. 6, p. 49, pl. 15, fig. 1,
1917; Carnegie Inst. Washington Publ. 311, p. 66, pl. 11, fig. 5, 1922;
U. S. Nat. Mus. Bull. 104, pt. 6, p. 31, pl. 3, figs. 7 a-c, 1929.—CusHMaN
and VALENTINE, Contr. Dept. Geol. Stanford Univ., vol. 1, no. 1, p. 12, pl. 3,
figs. 1 a—c, 1930.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 21
Miliolina costata Hrron-ALLEN and HARLAND, Trans. Zool. Soc. London, vol. 20,
pt. 2, p. 579, pl. 44, figs. 9-12, 1915.
Test elongate, two to three times as long as broad, periphery
usually rounded; chambers distinct, inflated; sutures somewhat de-
pressed ; wall ornamented by numerous fine longitudinal costae, often
slightly oblique to the axis of the chamber; aperture at the end of
a short neck, with a slight lip, and single short tooth. Length,
0.35-0.5 mm.; breadth, 0.15-0.25 mm.; thickness, 0.1-0.12 mm.
The types of d’Orbigny’s species are from the Mediterranean.
Forms such as those figured here occur rather abundantly in our
material from Vavau Anchorage, Tonga Islands, from which the
figured specimens came, and from the following Fiji Islands local-
ities: 12 fathoms off Levuka; 3 fathoms, Viva Anchorage; Mokaujar
Anchorage; and near Nairai. It was also common in 7 fathoms
at Rotonga.
QUINQUELOCULINA CRASSA @Orbigny var. SUBCUNEATA Cushman
PLATE 5, Ficures 8 a—c
Miliolina crassa HeroN-ALLEN and BARLAND (part) (not d’Orbigny), Trans.
Zool. Soc. London, vol. 20, pt. 2, p. 572, pl. 42, fig. 41 (not 37-40), 1915.
Quinqueloculina crassa D’ORBIGNY var. subcuneata CUSHMAN, U. 8. Nat. Mus.
Bull. 100, vol. 4, p. 4238, pl. 89, figs. 4 a—-c, 1921; Carnegie Inst. Washington
Publ. 342, p. 62, pl. 23, fig. 7, 1924; U. S. Nat. Mus. Bull. 104, pt. 6, p. 30,
pl. 5, figs. 1 a—c, 1929.
Quinqueloculina crassa CUSHMAN (not d’Orbigny), Carnegie Inst. Washington
Publ. 344, p. 82, 1926.
Test similar to the typical form in ornamentation but short, and
the chambers wedge-shaped, almost sharp at the peripheral angles.
This variety, originally described from the Philippines, occurs
widely distributed in the Indo-Pacific and also extends into the
Atlantic. ‘
The form is not common in our material, but occurs rarely off
Levuka, Fiji Islands, in 12 fathoms; Vavau Anchorage, Tonga
Islands, 18 fathoms; off Rotonga, 7 fathoms; Port Lotten, Kersail,
Caroline Islands; and Rongelap Atoll, Marshall Islands.
QUINQUELOCULINA CRENULATA, new species
PLATE 5, Figures 11 a—c
Test elongate, slender, about three times as long as broad ; chambers
distinct, not much inflated; sutures distinct; wall ornamented by
numerous very coarse, obliquely curved, short costae, extending in-
ward from the peripheral angle, and sloping toward the base of the
chamber, six at the base of the last-formed chamber sometimes form-
22 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
ing reticulations; aperture with a prominent neck. Length, 0.55-0.75
mm.; breadth, 0.18-0.18 mm.; thickness, 0.08-0.12 mm.
Holotype-—U.S.N.M. No. 14615, from Port Lotten, Kersail, Caro-
line Islands.
This is a very peculiarly ornamented species, and it somewhat re-
motely resembles the form described and named by d’Orbigny in his
Cuban papers as Quingueloculina tricarinata. Our species is, how-
ever, apparently quite distinct from d’Orbigny’s, and enough speci-
mens were found at the type locality to show that the characters are
fairly constant. It has not occurred, however, at any of the other
stations.
QUINQUELOCULINA FUNAFUTIENSIS (Chapman)
PLATE 5, Ficures 9, 10
Miliolina funafutiensis CHAPMAN, Journ. Linn. Soc. Zool., vol. 28, p. 178, pl. 19,
fig. 6, 1901; Proc. Zool. Soc. London, vol. 1, p. 231, 1902.—HERON-ALLEN and
EaRLanp(?), Trans. Zool. Soc. London, vol. 20, pt. 2, p. 566, pl. 42, figs. 21,
22, 1915.
Quinqueloculina funafutiensis CUSHMAN, Carnegie Inst. Washington Publ. 311,
p. 67, pl. 18, fig. 8, 1922; U. S. Nat. Mus. Bull. 104, pt. 6, p. 30, pl. 5,
figs. 1 a—c, 1929.
Test somewhat longer than broad in end view, with the angles
subacute or rounded, but generally triangular, occasionally with the
angles keeled; chambers distinct, somewhat inflated; sutures slightly
depressed; wall ornamented with numerous delicate costae which
are rather strongly oblique to the periphery of the chamber; aper-
ture at the end of a slight neck, rounded with a simple elongate
tooth. Length, 0.3-0.5 mm.; breadth, 0.18-0.25 mm.; thickness, 0.12—
0.2 mm.
Chapman originally described this species from the lagoon at
Funafuti in the Pacific; it has since been recorded from the Ke-
TABLE 4.—Quinqueloculina funafutiensis—material examined
W.
Num- Bot- |
U.S.N.M. | ber of| A!ba- : | tom | Character of | Occur-
No. speci- te Locality Depth Lec bottom 1 | rence
ture |
Baia ee eee
Fathoms | °F. |
21970._-_- 1 | H3849 | Village on west side Niau Atoll, AGT | ee co. s. pter. 0z_...| Rare.
1.75 miles NE.
| Do.
21971255 1 | H8866 | Lat. 17° 17’ S., long. 145° 45’ 30’” 804 | Wipe aa glob. oz. Mang--
1 Key to abbreviations is given in Table 1.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 23
rimba Archipelago off southeast Africa by Heron-Allen and Earland,
and I have had similar specimens from the Atlantic off the Tortugas,
which show the oblique costae.
Our material in the present collections shows that the species is
well distributed about the South Pacific Islands, occurring at Mo-
kaujar Anchorage, Fiji Islands; off Levuka, Fiji, in 12 fathoms;
Vavau Anchorage, Tonga Islands; both in the lagoon and off the
ished; aperture rounded or elliptical, with a distinct tooth, some-
times bifid at end. Length, 0.6 mm.; breadth, 0.35 mm.; thickness,
0.25 mm.
QUINQUELOCULINA cf. GRACILIS d’Orbigny
PLATE 5, Ficgurp 12
Our figure shows a very elongate specimen, which has a tapering
neck with a very distinct phialine lip and circular aperture. Such
forms occur at Rangiroa and are somewhat like d’Orbigny’s species,
although they seem to be distinct. There are slight traces of longi-
tudinal markings, and the wall is polished. Not enough specimens
were obtained to be sure of the complete characters.
QUINQUELOCULINA GUALTIERIANA d’Orbigny
PLATE 6, Ficures 1 a-c
Quinqueloculina gualtieriana D’ORBIGNY, in De la Sagra, Hist. Fis. Pol. Nat.
Cuba, “ Foraminiféres,” p. 186, pl. 11, figs. 1-3, 1839.
Test slightly longer than broad, with the periphery sharply angled
and the test tending to become more compressed as the later cham-
bers are added; chambers distinct, slightly inflated, broadest toward
the basal end, which projects somewhat beyond the general outline
of the test; sutures very distinct, depressed; wall smooth and _ pol-
ished; aperture rounded or elliptical, with a distinct tooth, sometimes
bifid at end. Length, 0.6 mm.; breadth, 0.35 mm.; thickness, 0.25 mm.
This species, described by d’Orbigny from the West Indian region,
is present in very typical form in our South Pacific material. In a
Jarge series there are a few specimens that seem to show that this
species is tending toward Massilina, as occasionally specimens are
found in which the last-formed chambers are in a single plane. The
whole test is much compressed, as shown in the original figures, and
the step to Masstlina would not be a very great one.
Our specimens are from off the Fiji Islands in 12 fathoms, Levuka;
3 fathoms, Viva Anchorage; in 7 fathoms off Rotonga; and most
abundant at Vavau Anchorage, Tonga Islands.
24 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
QUINQUELOCULINA LAMARCKIANA d’Orbigny
Puate 6, FIGURES 2 a-c
Quinqueloculina lamarckiana p’ORBIGNY, in De la Sagra, Hist. Fis. Pol. Nat.
Cuba, “ Foraminiféres,” p. 189, pl. 11, figs. 14, 15, 1839—CusHMaN, Proc.
U. S. Nat. Mus., vol. 59, p. 65, pl. 15, figs. 13, 15, 1921; Carnegie Inst.
Washington Publ. 311, p. 64, 1922; Publ. 342, p. 63, 1924; Publ. 344, p. 81,
1926.—HowsE, Journ. Pal., vol. 2, p. 175 (list), 1928——-CusHMAN, U. S. Nat.
Mus. Bull. 104, pt. 6, p. 26, pl. 2, figs. 6 a—c, 1929.—CuUSHMAN and VALENTINE,
Contr. Dept. Geol. Stanford Univ., vol. 1, no. 1, p. 10, pl. 1, figs. 9, 10,
1930.—CuUSHMAN and Moyer, Contr. Cushman Lab. Foram. Res., vol. 6, p.
52, 1930.—CusuMAN, Florida State Geol. Surv. Bull. 4, p. 20, pl. 2, figs. 3-5,
1930.
Quinqueloculina auberiana Dd’ OrBieny, in De la Sagra, Hist. Fis. Pol. Nat.
Cuba, “ Foraminiféres,” p. 193, pl. 12, figs. 1-3, 1839.
Quinqueloculina cuvieriana H. B, Brapy (not d’Orbigny), Rep. Voy. Challenger,
Zoology, vol. 9, p. 162, pl. 5, figs. 12 a—c, 1884.
Test nearly as broad as long; chambers distinct; sutures slightly
depressed; chambers generally triangular in transverse section, the
angles subacute but not carinate; wall smooth and polished; aper-
tural end of the chamber very slightly extended, forming an ellip-
tical neck without a definite lip, but with a narrow elongate tooth.
Maximum length, 1 mm.; breadth, 0.8 mm.; thickness, 0.5 mm.
This is a very widely distributed species occurring most commonly
in tropical waters, although there are numerous records for it in
comparatively deep water. Our specimens, in addition to the Adlba-
tross stations, data for which are given in the accompanying table,
are from off Fiji Islands, 40 to 50 fathoms; off Nairai, Fiji, 12
to 24 fathoms; off Levuka, Fiji, 12 fathoms; and Viva Anchorage,
Fiji, 3 fathoms. The species is common also at Vavau Anchorage,
Tonga Islands, in 18 fathoms.
TABLE 5.—Quinqueloculina lamarckiana—material examined
Num- Alba- ae
UEN-M. | bert ‘toss Locality Depth | tem. | | Charaetes of“) Ooo
mens Peres
ture
Fathoms | °F.
219612 1 | H8815 | Lat. 15° 15’ S., long. 147° 51’ 35’” O24 eee wh. co. s. brk. } Rare.
Ww. sh.
PAQG2. = 5 1 | H3827 | Lat. 14° 53’ 20’’ S., long. 148° 486). 25. ls ers. wh. co. Ss. Do.
42’ 30” W. vol. part.
21963 So 1 | H3873 | Southwest point Tahanae, 68° 966 |_----- glob. oz. mang__| Do.
N., 4 miles E.
21964_____- 1 | H3898 | Northwest point Hikueru Atoll, 348 | 43.8 | co. s. brk. sh.-.-| Do.
¥% mile E.
21965 sa 1 | H3914 | Northeast point Nukutavake, 636; 88.9)! cos S#22--. 2222554 Do.
1 mile 8.
2196622 25 1 | H3935 | Hereheretue Atoll, 1 mile W---- 594 | 39.5 | crs. co, S_-.---2-- Do.
2196 eee a 2 | H3936 | Hereheretue Atoll, 0.3 mile E___- 189 | 62.1 | co.s.mang. part_| Do.
1 Key to abbreviations.is given in Table 1.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 25
The species was originally described under the first two names by
d’ Orbigny from the West Indies, where it is well distributed. Many
of the West Indian species occur in our Indo-Pacific material in
rather typical form.
QUINQUELOCULINA PARKERI (H. B. Brady)
PLATE 6, FIGURES 3, 4
«“Quinqueloculina with oblique ridges ” PARKER, Trans. Micr. Soc. London, vol. 6,
p. 53, pl. 5, fig. 10, 1858.
Miliolina parkeri H. B. Brapy, Quart. Journ. Micr. Sci., vol. 21, p. 46, 1881;
Rep. Voy. Challenger, Zoology, vol. 9, p. 177, pl. 7, fig. 14, 1884—-CHAPMAN,
Proc. Zool. Soe. London, 1895, p. 11; Journ. Linn. Soe. Zool., vol. 28, p. 175,
1901; Proce. Zool. Soe, London, vol. 1, p. 231, 1902.—DAKIN, Rep. Pearl
Oyster Fisheries Ceylon, p. 230, 1906.—HERON-ALLEN and HARLAND, Trans.
Zool. Soe. London, vol. 20, pt. 2, p. 574, pl. 48, figs. 11, 12, 1915.
Quinqueloculina parkeri CUSHMAN, U. S. Nat. Mus. Bull. 71, pt. 6, p. 50,
pl. 15, fig. 83, 1917; Carnegie Inst. Washington Publ. 213, p. 290, 1918; U. S.
Nat. Mus. Bull. 100, vol. 4, p. 440, pl. 86, figs. 4 a—c, 1921; Carnegie Inst.
Washington Publ. 342, p. 59, pl. 22, fig. 3, 1924.
Test about twice as long as broad, periphery subacute; chambers
distinct; sutures depressed; surface marked by a series of transverse
ridges and alternating excavations not involving the peripheral mar-
gin; aperture with a slight lip, rectangular, with a single tooth.
Length, 1-1.65 mm.; breadth, 0.6-1 mm.; thickness, 0.4-0.6 mm.
This typically Indo-Pacific species, which from published records
has a wide range, occurs rather sparingly in our collections, although
it has occurred in few numbers at the following stations: Off the Fiji
Islands in 12 fathoms at Levuka, and at Mokaujar Anchorage; in
18 fathoms, Vavau Anchorage, Tonga Islands; Makemo Lagoon,
Paumotu Archipelago; Port Lotten, Kersail, Caroline Islands; and
Guam Anchorage, Ladrone Islands, in 21 fathoms. Specimens from
the last locality are the largest and best developed of all these, and
the figured specimens are from there.
QUINQUELOCULINA POLYGONA d’Orbigny
PLATE 6, FIGURES 5, 6
Quinqueloculina polygona D’ORBIGNY, in De la Sagra, Hist. Fis. Pol. Nat. Cuba,
* Foraminiféres,” p. 198, pl. 12, figs. 21-23, 1889.—CusHMAN, Proc. U. S. Nat.
Mus., vol. 59, p. 66, pl. 16, figs. 3, 4, 1921; Carnegie Inst. Washington Publ.
311, p. 68, 1922; Publ. 344, p. 82, 1926; U. 8S, Nat. Mus. Bull. 104, pt. 6, p. 28,
pl. 3, figs. 5 a—e, 1929.
Test two or three times as long as broad; chambers distinct, each
end extended beyond the previous outline, polygonal in transverse
section with the periphery either truncate and the angles somewhat
rounded, or with definite projecting carinae at the angles; sutures
26 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
very distinct, usually slightly depressed; wall either smooth or more
frequently with slight elongate pits covering much of the surface;
aperture elongate, somewhat rectangular, but usually narrowest
toward the base, toward each a simple narrow tooth extends into the
opening. Length, 1-1.3 mm.; breadth, 0.4-0.5 mm.; thickness, 0.2—
0.25 mm.
D’Orbigny described this species originally from the West Indies,
from Jamaica and Cuba. The Indo-Pacific specimens seem to show
a wide variation the extremes of which seem very distinct, but with
a large series these distinctions are usually bridged.
It is one of the commonest species in our collections, occurring off
Fiji Islands at Mokaujar Anchorage; 12 fathoms off Levuka; Vavau
Anchorage, Tonga Islands; Rutavu; Makemo Lagoon; Pinaki; and
beach off wharf Hereheretue. There are also specimens noted in the
accompanying table from two Albatross stations.
TABLE 6.—Quinqueloculina polygona—maierial examined
Bot-
Num-
Alba- tom
U.S.N.M. | ber of : Character of Occur-
No. speci- atabne Locality Depth | tem- bottom 1 rence
mens Pelee
ture
Fathoms| ° F.
21968 - 22-<e 2 | H3916 | Pinaki Atoll, 1 mile E_--_-____-- 486 41 | ers.co.s. pter.oz| Rare.
21969202. 1 | H3924 | Nukutipipi Atoll, 1 mile NW--- 649 39\| co.8. DEK: Sh2=22| ) Do.
1 Key to abbreviations is given in Table 1.
QUINQUELOCULINA SAMOAENSIS Cushman
PLATE 7, F1IGuRES 1 a—c
Quinqueloculina samoaensis CUSHMAN, Carnegie Inst. Washington Publ. 342,
p. 59, pl. 21, figs. 4-7, 1924.
Test slightly longer than broad, periphery either squarish in the
adult or with a blunt keel in the young, often sinuous; wall thick,
with an encrusting of fine sand grains, initial end of the chamber
extended and the apertural end elongated slightly into a cylindrical
neck with a phialine lip and single tooth. Length, 0.65 mm.;
breadth, 0.45 mm.; thickness, 0.3 mm.
This species, originally described from shallow water collections
at Samoa, seems to be widely distributed in the collections examined,
occurring at the following localities: Mokaujar Anchorage, Fiji;
Levuka, Fiji, 12 fathoms; off Nairai, Fiji, 12-24 fathoms; inside
the lagoon, Pinaki Atoll, Paumotus; off Rotonga, 7 fathoms; Vavau
Anchorage, Tonga Islands, 18 fathoms; off Niau; Port Lotten,
Kersail, Caroline Islands; and Guam Anchorage, Ladrone Islands,
21 fathoms.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 27
There is considerable variation in the sinuous character of the
chambers, but except for this variation the other characters seem to
be generally constant. It may be easily distinguished from either
of the other two arenaceous forms which are common in the region.
QUINQUELOCULINA SEMIRETICULOSA, new species
PLATE 7, FIGURES 2 a, Db
Test small, two to three times as long as broad, periphery rounded,
the basal end broadly rounded, the apertural end somewhat more
contracted; chambers distinct, slightly inflated; sutures slightly
depressed; wall beautifully ornamented by a somewhat reticulate
pattern, the main elements of which are composed of oblique costae,
somewhat irregularly sinuous, the depressed areas between broken
up by transverse ridges into a series of elongate pits; aperture very
slightly produced, occasionally with a very short neck, the aperture
itself nearly circular with a very short, simple tooth. Length,
0.4-0.5 mm.; breadth, 0.2-0.25 mm.; thickness, 0.15-0.2 mm.
Holotype—Cushman Coll. No. 14619, from Port Lotten, Kersail,
Caroline Islands.
This seems to be a very beautifully ornamented, localized species,
which was fairly common at the type locality but was not seen in
any of the other collections.
QUINQUELOCULINA STRIATULA, new species
PLATE 7, Ficures 3, 4
Test slightly longer than broad, the periphery of the chambers
subacute; chambers triangular in section, distinct; sutures distinct,
only slightly depressed; wall ornamented with numerous very fine,
incised lines, slightly oblique to the periphery of the chamber, the
peripheral angle of the chamber usually smooth; aperture large,
broad, with a very slight lip, and a large, fiat tooth nearly filling
the aperture. Length, 0.6-0.8 mm.; breadth, 0.55-0.65 mm.; thick-
ness, 0.35—0.4 mm.
Holotype—Cushman Coll. No. 14780, from Mokaujar Anchorage,
Fiji Islands.
This is abundant at the type locality, and also occurs about Fiji
from Viva Anchorage, 3 fathoms; off Levuka, 12 fathoms; off Nairai,
12 fathoms. It is evidently widely distributed in the South Pacific,
as we have less abundant specimens from Rotonga, 7 fathoms;
Pinaki Islands; Makemo Lagoon; and Rongelap Atoll, Marshall
Islands.
This is evidently the same as Afiliolina undulata of Heron-Allen
and Earland, not of d’Orbigny, figured in their Kerimba paper
91513—32——3
28 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
(pl. 48, figs. 5-8).2 A reference to the figures of Quingueloculina
undulata d’Orbigny given by Fornasini shows that the species has
a rather narrow, elongate aperture, and a very narrow tooth. Speci-
mens in my collection from Castel Arquato, Italy, one of the locali-
ties mentioned by d’Orbigny, are entirely identical with the original
figures of d’Orbigny and show the same type of aperture and general
characters of the test, the outer angles of which are not nearly so
sharply formed as in our South Pacific species. The figures given
by Heron-Allen and Earland show specimens with the same broad
aperture and flat tooth that are seen throughout our series. They
~ mention the close relation of their material to Quinqueloculina
nussdorfensis d’Orbigny, but topotype material of that species com-
pared with the figures shows that it has a narrow tooth and an elon-
gate aperture, with the sides truncate. They also mention 7'rélocu-
lina brongniartiana d’Orbigny in this connection, but that species
as developed in the West Indies is triloculine with a rounded periph-
ery, and an exserted neck, making it a very different species from
ours.
The material that I have referred to Triloculina suborbicularis
d’Orbigny * is also possibly to be referred to this same species.
QUINQUELOCULINA SULCATA d’Orbigny
PLATE 7, FIGURES 5-8
Quinqueloculina sulcata p’OrBIeNy, Ann. Sci. Nat., vol. 7, p. 801, no. 17, 1826.—
ForNASINI, Mem. Accad. Sci. Istit. Bologna, ser. 5*, vol. 8, p. 368, fig. 9 (in
text), 1900.
Test elongate, 214 to 3 times as long as broad, apertural end con-
siderably extended out beyond the main body of the test; chambers
distinct, elongate, in the early stages with a single angle at the
periphery, later becoming truncate with two distinct angles, and in
the adult typically with three raised costae; sutures fairly distinct,
not much depressed; wall smooth except for the costae; both ends
of the last-formed chamber extending beyond the previous chambers,
the apertural end tapering with a rounded opening, a definite lip,
and a simple tooth, which extends slightly beyond the rim of the
aperture in side view. Maximum length, 1.75 mm.; breadth, 0.6
mm.; thickness, 0.8 mm.
D’Orbigny’s original material of this species was from the Red
Sea, an area that is closely associated in its fauna with that of the
general Indo-Pacific region. Although this species was not really
known until the publication of Fornasini’s paper in 1900, it seems
2 The Foraminifera of the Kerimba Archipelago, Trans. Zool. Soc. London, vol. 20, pt. 2,
pp. 5438-794, pls. 40-53, 1915.
$U. S- Nat. Mus. Bull. 71, pt: 6, pl. 21, fig. 3, 1917.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 29
to fit this particular species much better than any of the earlier fig-
ures. The development as shown in our series of specimens is inter-
esting. Though the neck remains of the same general character
throughout and also the general form of the test, the form of the
chambers changes greatly from the young to the adult condition, as
already noted in the above description.
This is one of the largest species of the genus in the collection and
is best developed in the collections from Guam Anchorage, Ladrone
Islands, in 21 fathoms. It also occurs off the Fiji Islands at Mokaujar
Anchorage and at Rangiroa. There are numerous specimens from
various localities that are possibly the young of this species, but
where no adults are present to check this they have not been included.
Genus SCHLUMBERGERINA Munier-Chalmas, 1882
Schlumbergerina MUNIER-CHALMAS, Bull. Soc. Géol. France, ser. 3, vol. 10,
p. 424, 1882.—CusHMAN, Cushman Lab. Foram. Res. Spec. Publ. No. 1,
p. 148, 1928; U. S. Nat. Mus. Bull. 104, pt. 6, p. 35, 1929.
Miliolina (part) of authors.
Massilina (part) of authors.
Genoholotype—Schlumbergerina areniphora Munier-Chalmas.
Test typically quinqueloculine, the chambers narrowing so that
frequently more than five chambers may be visible from the exterior;
wall calcareous, imperforate, the exterior thickly coated with sand
grains; aperture cribrate.
Late Tertiary and Recent.
The Miliolina alveoliniformis H. B. Brady described in 1879 is
probably the same as Munier-Chalmas’s species, and belongs here.
This species is often abundant in shallow-water tropical collections,
especially in the Indo-Pacific, but similar forms are also found in
shallow warm water in the West Indian region.
SCHLUMBERGERINA ALVEOLINIFORMIS (H. B. Brady)
PLATE 8, FicurEe 1
Miliolina alveoliniformis H. B. Brapy, Quart. Journ. Micr. Sci., vol. 19, p. 268,
1879; Rep. Voy. Challenger, Zoology, vol. 9, p. 181, pl. 8, figs. 15-20, 1884.—
Eccer, Abh. kén. bay. Akad. Wiss. Miinchen, Cl. II, vol. 18, p. 232, pl. 2,
figs. 17-19, 1893—Woopwarp, The Observer, vol. 4, p. 76, 1893.—MILLETT
(in part), Journ. Roy. Micr. Soc., 1898, p. 510—CHApMAN, Journ. Linn.
Soe. Zool., vol. 28, p. 177, 1900; vol. 30, p. 398, 1910.—Hrron-ALLEN and
EARLAND, Trans. Zool. Soc. London, vol. 20, pt. 2, p. 581, 1915.
Quinqueloculina alveoliniformis CUSHMAN, U. S. Nat. Mus. Bull. 71, pt. 6, p. 43,
1917; U. S. Nat. Mus. Bull. 100, vol. 4, p. 443, 1921; Proc. U. S. Nat. Mus.,
vol. 59, p. 64, 1921; Carnegie Inst. Washington Publ. 311, p. 64, 1922;
Publ. 342, p. 58, pl. 21, fig. 8, 1924; Publ. 344, p. 81, 1926.
Schlumbergerina alveoliniformis CusHMAN, U. S. Nat. Mus. Bull. 104, pt. 6,
p. 36, 1929.
30 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Test elongate, fusiform, composed of numerous chambers, long and
narrow, five normally visible from the exterior in the early stages,
more in the adult; wall in young specimens thin and porcelaneous,
in adults covered with sand grains; aperture composed of numerous
pores, or radiate, typically cribrate. Length, 1.5-2.2 mm.; breadth,
0.8-1 mm.
The species is abundant in the Fiji Islands at Mokaujar Anchorage,
but it has not occurred in the other shallow-water collections. There
is a specimen from Albatross Station H3875, half a mile off south-
west point of Tahanae, Paumotus, in 269 fathoms. The bottom char-
acter at this station is given as coarse coral sand, and this is un-
doubtedly a specimen washed out from shallow water. It is sur-
prising that this species did not occur at more localities, as it is
widely distributed in the Indo-Pacific.
Genus MASSILINA Schlumberger, 1893
Massilina SCHLUMBERGER, Mém. Soc. Zool. France, vol. 6, p. 218, 1893.—-CusHMAN,
Cushman Lab. Foram. Res. Spec. Publ. No. 1, p. 149, 1928; U. S. Nat. Mus.
Bull. 104, pt. 6, p. 36, 1929.
Quinqueloculina (part) p’OrBieny, Ann. Sci. Nat., vol. 7, p. 303, 1826.
Miliolina (part) of authors.
Genotype.—By designation, Quinqueloculina secans d’Orbigny.
Test with the early chambers quinqueloculine, later ones added on
opposite sides in a single plane, the quinqueloculine stage present
in both megalospheric and microspheric forms; aperture simple, with
a bifid tooth.
Some of our specimens show very clearly the development of this
genus from Quingueloculina, the early stages being decidedly quin-
queloculine, while the later ones progressively add the chambers
more and more in a single plane. The genus may be distinguished
from Spiroloculina, which has almost all the chambers in a single
plane from the beginning, especially in the megalospheric form. In
Massilina all the early chambers are quinqueloculine in both the
microspheric and megalospheric forms. Species occur in which the
exterior is arenaceous, but most of the species have lost that character
and are entirely calcareous.
MASSILINA ALVEOLINIFORMIS Millett
PLATE 8, FIGURES 3 a, 0D
Massilina alwveoliniformis Mitierr, Journ. Roy. Micr. Soc., 1898, p. 609, pl. 13,
figs. 5-7.—HERON-ALLEN and Hartanp, Trans. Zool. Soe., London, vol. 20, pt.
2, p. 584, pl. 45, fig. 15, 1915.—CusHMAN, Carnegie Inst. Washington Publ.
311, p. 69, 1922; Publ. 342, p. 64, 1924; U. S. Nat. Mus. Bull. 104, pt. 6, p. 39,
1929.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS Sl
Spiroloculina asperula H. B. Brapy (not Karrer), Rep. Voy. Challenger,
Zoology, vol. 9, p. 152, pl. 8, figs. 18, 14 (117), 1884—CusHMAN, Proc.
U. S. Nat. Mus., vol. 59, p. 72, 1921.
Massilina asperula CUSHMAN, U. S. Nat. Mus. Bull. 100, vol. 4, p. 447, 1921.
Quinqueloculina variabilis p’OrBIaNy, Ann. Sci. Nat., vol. 7, p. 301, No. 15,
1826.—ForNASINI, Mem. Accad. Sci. Istit. Bologna, ser. 6", vol. 2, p. 65,
pl. 3, figs. 6, 7, 1905.
Test much compressed, nearly circular; early chambers quinque-
loculine, later ones in a single plane, periphery rounded, chambers
distinct; wall of sand grains with a porcelaneous lining to the cham-
bers; aperture very slightly exserted. Length, 1 mm.; breadth, 0.9
mm.; thickness, 0.25 mm.
This species, except for a very few records in the Tortugas region
off Florida, is known only from the Indo-Pacific, but it is apparently
widely distributed in that region. The only material we have in the
present collection that can be referred to it is from off Levuka, Fiji
Islands, 12 fathoms.
It may be noted here that the Quinqueloculina variabilis of
d’Orbigny’s 1826 paper, which came from Rawack, is probably the
same as Millett’s species. As the name Quinqueloculina variabilis
d’Orbigny remained a nomen nudum until the publication of Forna-
sini in 1905, d’Orbigny’s name can not be used for this species.
MASSILINA PLANATA, new species
PxLaTE 8, Fiaures 8 a, 6
Test comparatively large, very much compressed, periphery
subacute, both ends of the final chamber projecting beyond the body
of the test; chambers distinct, much compressed, early ones entirely
quinqueloculine, later ones in a single plane; sutures fairly distinct,
but very slightly depressed; wall matte, smooth except for very fine
elongate pits arranged generally in the long axis of the chamber;
aperture elliptical, with a slight lp and an elongate simple tooth,
shghtly bifid at the tip. Length of holotype, 2 mm.; breadth, 1 mm.;
thickness, 0.2 mm.
Holotype—Cushman Coll. No. 14629, from Guam Anchorage,
Ladrone Islands, 21 fathoms.
This is a large striking species that in our material occurred at
the type locality only. It suggests rather strongly the specimens re-
ferred by Heron-Allen and Earland in their Kerimba Archipelago
paper (pl. 41, figs. 1-5)* to Spiroloculina planissima (Lamarck).
The early quinqueloculine chambers show that their species belongs
to the genus Masstlina and not to Spiroloculina, and its general char-
acter suggests that our species is to be found widely distributed in the
Indo-Pacific region.
4The Foraminifera of the Kerimba Archipelago, Trans. Zool. Soc. London, vol. 20, pt. 2,
pp. 543-794, pls. 40-53, 1915.
32 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
MASSILINA INAEQUALIS Cushman
PLATE 8, FIcuREs 6, 7
Massilina inaequalis CUSHMAN, Proc. U. S. Nat. Mus., vol. 59, p. 72, pl. 17,
figs. 12, 18, 1921; U. S. Nat. Mus. Bull. 104, pt. 6, p. 38, pl. 7, figs. 6 a—c, 1929.
Test longer than broad, compressed, periphery squarely truncate;
chambers elongate, of uniform width, quadrangular in transverse
section; sutures distinct, very slightly depressed; wall smooth except
for numerous fine, linear depressions breaking up the evenness of
the surface; aperture rounded, with a sight lip and a definite linear
tooth which is slightly bifid at the tip. Length, 1.3 mm.; breadth,
0.6 mm.; thickness, 0.18 mm.
This species was originally described from the north coast of
Jamaica at Montego Bay, where it was fairly common. It has not
been recorded elsewhere, but very typical specimens occur in our
material from Vavau Anchorage, Tonga Islands, which seem to be
absolutely identical with the Jamaican specimens.
MASSILINA AUSTRALIS, new species
PLATE 8, FIGURES 2 a, 0D
Test very much compressed, the earliest chambers quinqueloculine,
later ones becoming planispiral, with slightly more than two cham-
bers making up a coil, periphery subacute; chambers distinct, very
slightly inflated; sutures fairly distinct, very slightly depressed be-
tween the last-formed chambers, but those of the earlier portion
obscured ; wall fairly thin, usually translucent, finely pitted; aperture
elongate, narrow, with a slight lip, but no apertural tooth. Diameter,
0.8-1 mm.; thickness, 0.1 mm.
Holotype—U.S.N.M. No. 14625, from Rotonga (=? Rarotonga,
Cook Islands), 7 fathoms.
This species is very common at the type locality. It is probably
the same as that assigned by some authors to Massilina secans of
d’Orbigny. That species, however, in its typical development has a
very prominent quinqueloculine stage, and the later chambers have
a definite elongate tooth in the aperture. Our series of specimens
from the type locality shows that some of the last-formed chambers
have a wrinkled appearance as if due to stages of growth of the
chamber. The surface is dull and seems to be slightly pitted all
over.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 33
MASSILINA MACILENTA (H. B. Brady)
PuatE 8, Fiaures 4 a, b
Miliolina macilenta H. B. Brapy, Rep. Voy. Challenger, Zoology, vol. 9, p.
167, pl. 7, figs. 5, 6, 1884——CHaArmMAn, Journ. Linn. Soc. Zool., vol. 28, p.
399, 1902.
Quinqueloculina macilenta CUSHMAN, U. S. Nat. Mus. Bull. 71, pt. 6, p. 55,
1917.
(?)Massilina macilenta Hrron-ALLEN and HarLAnp, Trans. Zool. Soc. London,
vol. 20, p. 583, pl. 45, figs. 18, 14, 1915—CusHMaN, Carnegie Inst.
Washington Publ. 342, p. 64, pl. 24, figs. 3, 4, 1924.
(?)Massilina secans D’ORBIGNY var. macilenta MILLETT, Journ. Roy. Micr. Soc.,
1898, p. 609, pl. 18, fig. 4.
Test nearly circular, much compressed, periphery rounded, early
chambers quinqueloculine, later ones spiroloculine; surface orna-
mented by obliquely longitudinal costae; apertural end with a flar-
ing lip, aperture elongate. Length, 0.35 mm.; breadth, 0.8 mm.;
thickness, 0.1 mm.
The records for this species show that it is rather definitely limited
to the Indo-Pacific regions. It is a very distinctive, although small
species, and shows little variation in its characters. Specimens are
rare in our material, but represent the following localities: Off Fiji
Islands in 12 fathoms, Levuka, and Mokaujar Anchorage; near
Niau, and Vavau Anchorage, Tonga Islands, 18 fathoms.
MASSILINA CRENATA (Karrer)
PLATE 8, FIGURES 5 a, b
Spiroloculina crenata Karrer, Sitz. Akad. Wiss. Wien, vol. 57, p. 185, pl. 1,
fig. 9, 1868.—H. B. Brapy, Rep. Voy. Challenger, Zoology, vol. 9, p. 156, pl.
10, figs. 24-26, 1884.
Massilina crenata CUSHMAN, U. S. Nat. Mus. Bull. 71, pt. 6, p. 57, pl. 20, fig 2,
1917; U. S. Nat. Mus. Bull. 100, vol. 4, p. 455, 1921; Carnegie Inst. Wash-
ington Publ. 311, p. 69, pl. 11, fig. 6, 1922; Publ. 342, p. 66, pl. 25, fig. 4,
1924; U. S. Nat. Mus. Bull. 104, pt. 6, p. 38, pl. 7, fig. 5, 1929.
Test in its early development quinqueloculine, adult chambers
in a single plane, in front view subelliptical, nearly as broad as long,
compressed; chambers long and narrow, margin crenulate, due to
the regular contractions or plications of the chamber in the adult;
aperture rounded. Length, 0.55 mm.; breadth, 0.35 mm.; thickness,
0.08 mm.
This is typically an Indo-Pacific species, but there are rare speci-
mens from the West Indian region.
In the present collections the species has occurred at but two
stations, at Rangiroa and at Albatross Station H3858, 28° south, 3.5
miles east of Ngaruae Pass, Fakarava Atoll, Paumotus. A specimen
from the latter station is figured.
34 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Genus SPIROLOCULINA d’Orbigny, 1826
Spiroloculina D’Orpieny, Ann. Sci. Nat., vol. 7, p. 298, 1826——H. B. Brapy,
Rep. Voy. Challenger, Zoology, vol. 9, p. 147, 1884.—-CHAPMAN, The Foram-
inifera, p. 89, 1902——CusHmMAN, Cushman Lab. Foram. Res. Spec. Publ.
No. 1, p. 149, 1928; U. S. Nat. Mus. Bull. 104, pt. 6, p. 40, 1929.
Miliola (part) Lamarck, Ann. Mus. d’Hist. Nat., vol. 5, p. 852, 1804 (and later
authors).
Genotype.—By designation, Spiroloculina depressa d’Orbigny.
Test with the early chambers in the microspheric form quinquelo-
culine, the later ones in a single plane; chambers a half coil in
length, in the megalospheric form all the chambers usually in one
plane; apertural end usually with a neck and lip, simple, with a
simple or bifid tooth.
Permian (?) Cretaceous to Recent.
This is a more specialized genus than Masstlina, and the character
of having chambers in a single plane is taken on very early even
in the microspheric form. In the shallow-water material of the
Pacific islands, specimens of this genus are often very abundant but
belong to but few species.
SPIROLOCULINA GRATELOUPI d’Orbigny
PLATE 8, Figures 9-11
Spiroloculina grateloupi p’OrBieny, Ann. Sci. Nat., vol. 7, p. 298, 1826.—TERQquEM,
Mém. Soc. Géol. France, ser. 3, vol. 1, p. 52, pl. 5, figs. 5, 6, 1878.—WHdIESNER,
Arch. Prot., vol. 25, p. 208, 1912—-CusHMan, U. S. Nat. Mus. Bull. 71, pt. 6,
p. 31, pl. 4, figs. 4, 5, 1917; Proc. U. S. Nat. Mus., vol. 56, p. 634, 1919; U. S.
Nat. Mus. Bull. 100, vol. 4, p. 396, pl. 78, figs. 4 a, b; pl. 100, fig. 3; figs. 17, 18
(in text), 1921; Carnegie Inst. Washington Publ. 311, p. 59, 1922; Publ. 342,
p. 56, pl. 20, figs. 3, 4, 1924; Publ. 344, p. 80, 1926; U. S. Nat. Mus. Bull. 104,
pt. 6, p. 40, pl. 8, figs. 1 a, b, 1929.
Spiroloculina excavata H. B. Brapy (not d’Orbigny), Rep. Voy. Chalienger,
Zoology, vol. 9, p. 151, pl. 9, figs. 5, 6, 1884.
Spiroloculina impressa H. B. Brapy (not Terquem), Rep. Voy. Challenger, Zool-
ogy, vol. 9, p. 151, pl. 10, figs. 3, 4, 1884.
Spiroloculina angulosa p’Orpieny, Ann. Sci. Nat., vol. 7, p. 298, 1826.—ForNasINI,
Mem. Accad. Sci. Istit. Bologna, ser. 6*, vol. 1, p. 5, pl. 1, fig. 8, 1904.
Test elongate, broadest in the center, tapering toward either end;
chambers rapidly thickening as added, in end view the periphery
much the broadest portion of the test, central portion deeply exca-
vated ; periphery of the chambers in end view much convex, especially
in the central portion, the edges broadly rounded; chambers evenly
curved, the final chamber somewhat projecting, both at the base and
at the apertural ends, the latter having a decided neck with a phialine
lip; the aperture itself rounded and with either a single tooth with a
bifid end, the two projections forming a concave extremity, or in some
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 35
cases a pair of such bifid teeth opposite one another; surface of the
test dull, somewhat roughened. Length, 1.25 mm.; breadth, 0.6 mm. ;
thickness, 0.25 mm.
The typical form of the species is extremely common at some of the
South Pacific stations, especially about the Fiji Islands, where it
occurs off Levuka, 12 fathoms; Nairai, 12 fathoms; Viva Anchorage,
3 fathoms; and Mokaujar Anchorage; as well as at a station locality
not given, but in 40 to 50 fathoms. Besides these Fiji stations 1t
occurs at Vavau Anchorage, Tonga Islands; off Rotonga, 7 fathoms;
at Port Lotten, Kersail, Caroline Islands; and at Guam Anchorage,
Ladrone Islands, 21 fathoms. In the Albatross samples it occurs at
Stations H3848, off Point Venus, Tahiti Island, 807 fathoms; H3930,
off Anu Anuraro Atoll, Paumotus, 438 fathoms; and H3992, Schisch-
marev Pass, Marshall Islands, 482 fathoms.
TABLE 7.—Spiroloculina grateloupi—material examined
Num alba thn
U.S.N.M. | ber of % : Character of Occur-
No speci- AS es Locality Depth | tem- bottom ! rence
Pinan ion pera-
| ture
|
Fathoms| ° F.
21052 22 ar 1 | H3843 | Point Venus, Tahiti Island, 55° 16 Bea ee fne. vol. s. m____| Rare.
S., 3.8 miles E.
21955-= . 23! 1 | H3930 | Anu Anuraro Atoll, 34 mile N W_| ASSr| 40761 COS! = — ee _ -2e Do.
21956...---| 1 | H3992 | Schischmarev Pass, Wotje,Mar-| 482 | 41.7 |___-- dase ues! Do.
shall Islands, 1 mile N. |
1Key to abbreviations is given in Table 1.
SPIROLOCULINA GRATELOUPI d’Orbigny SERRULATA, new variety
PLATE 9, F1GuRES 1 a, D
Variety differing from the typical in the peripheral margins which
in the variety are beautifully serrate with a series of irregularly
shaped teeth.
Holotype of variety—Cushman Coll. No. 14748, from Viva An-
chorage, Fiji Islands, in 3 fathoms.
The figured specimen shows the holotype, which has the teeth very
well developed along the edges. There seem to be many gradations
between this and the form in which these edges are only roughened.
The variety was only found at the type locality, but there it is
abundant.
SPIROLOCULINA GRATELOUPI d’Orbigny ACESCATA, new variety
PLATE 9, FIGURES 2 a, Db
Spiroloculina canaliculata CUSHMAN (not d’Orbigny), Carnegie Inst. Wash-
ington Publ. 342, p. 57, pl. 21, fig. 1, 1924.
36 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Variety with the test elongate, elliptical, much compressed, some-
what variably depressed in the middle; chambers distinct, earlier
ones rectangular in transverse section, periphery truncate, the angles
sharply keeled, later chambers much compressed with a single keel;
sutures distinct, little depressed; wall smooth, glossy; apertural end
extended into a cylindrical neck and a distinct phialine lip and tooth.
Holotype of variety —Cushman Coll. No. 14747, from off Nairai,
Fiji Islands, 24 fathoms.
This variety is very abundant at the type locality, and there are a
few other records for it: Mokaujar Anchorage, Fiji; Levuka, Fiji.
T have already noted this variety from Samoa. The young stages,
which are found with the adults, are very similar to Spzroloculina
grateloupi, but as the form develops toward its adult stage there
is a very considerable compression of the test, and, in the adult,
chambers are developed that are not only compressed but have a
definite thin peripheral keel.
SPIROLOCULINA ANTILLARUM d’Orbigny
PLATE 9, FiacurEs 3-5
Spiroloculina antillarum p’OrBIaNy, in De la Sagra, Hist. Fis. Pol. Nat. Cuba,
“ Foraminiféres,” p. 166, pl. 9, figs. 3, 4, 1889.—H. B. Brapy, Rep. Voy.
Challenger, Zoology, vol. 9, p. 155, pl. 10, figs. 21 a, b, 1884——CusHMAN,
U. S. Nat. Mus. Bull. 100, vol. 4, p. 407, pl. 81, figs. 4 a, b; pl. 83, fig. 4 (7),
1921; Proc. U. 8S. Nat. Mus., vol. 59, p. 63, pl. 14, figs. 14, 15, 1921; Car-
negie Inst. Washington Publ. 311, p. 61, 1922; Publ. 342, p. 55, pl. 20,
fig. 1, 1924; U. S. Nat. Mus. Bull. 104, pt. 6, p. 43, pl. 9, fig. 3, 1929.
Test elongate, elliptical; chambers nearly circular in cross section;
surface ornamented by numerous longitudinal costae, often slightly
oblique, both ends of the chamber projecting and the intermediate
portions thus left either filled by a plate of clear material or occa-
sionally open; apertural end projecting and forming a cylindrical
neck with a slight lip and a single tooth, sometimes bifid at the
tip. Length, 0.55-0.75 mm.; breadth, 0.35-0.4 mm.; thickness,
0.15-0.18 mm.
The original specimens of this species came from off the West
Indies and were described by d’Orbigny in 1839. His species was
allowed to lapse, and much of the material that should have been
recorded as his species has been known under the name of S. grata
Terquem, because of the adoption of that name by Brady in the
Challenger report for this species of d’Orbigny. As a rule the typi-
cal form is more involute than the following variety (angulata),
and in the Pacific at least it is much less common. The chambers
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS aT
are circular in the transverse section in the typical form. Typical
specimens occur at the following localities: Off Nairai, Fiji, 24
fathoms; Rutavu; Rangiroa; and at the following Albatross Sta-
tions: H3853, Pakaka entrance to Apataki Lagoon, Paumotus, 613
fathoms; H3910, 1 mile off southwest point of Aki Aki, Paumotus,
877 fathoms; H3916, 1 mile off Pinaki Atoll, Paumotus, 486 fathoms;
H3983, half a mile off entrance of South Pass, Rongelap Atoll,
Marshall Islands, 400 fathoms.
TABLE 8.—Spiroloculina antillarum—nvtterial examined
U.S.N.M heel Alba- ‘ tom Character of Occur-
“No. | speci- eos Locality Depth ae bottom ! rence
mens reer
Fathoms | ° F.
21h Teese 2) 1 | H3853 | Pakaka entrance to Apataki BLS S924. | eos ivol =. 252 Rare.
Lagoon, 50° N., 2 miles E__--_
21968=525—- 1 | H3910 | Southwest point Aki Aki, 1 377 | 43 CO 5Sise. es 523: Do.
mile E.
219502 Fe 1 | H3916 | Pinaki Atoll, 1 mile E---.-_-~-- 486 | 41 crs.co.s. pter.oz.| Do.
219602222 = 2 | H3983 | Entrance to South Pass, Ronge- AQOW A804 I COs See nae tenes Do.
lap, Marshall Islands, }4 mile
N.
1 Key to abbreviations is given in Table 1.
SPIROLOCULINA ANTILLARUM d’Orbigny var. ANGULATA Cushman
PLATE 9, FIGURES 6-9:
Spiroloculina grata H. B. Brapy (part) (not Terquem), Rep. Voy. Challenger,
Zoology, vol. 9, pl. 10, figs. 22, 23, 1884.
Spiroloculina grata TERQUEM var. angulata CUSHMAN, U. S. Nat. Mus. Bull. 71,
pt. 6, p: 36, pl. 7, fig. 5, 1917.
Spiroloculina antillarwn D’ORBIGNY var. angulata CusuMAN, U. S. Nat. Mus.
Bull. 100, vol. 4, p. 408, pl. 81, figs. 5 a, b, 1921; Carnegie Inst. Washington
Publ. 311, p. 62, 1922; Publ. 342, p. 55, pl. 20, fig. 2, 1924; Publ. 344, p. 80,
1926; U. S. Nat. Mus. Bull. 104, pt. 6, p. 48, pl. 9, figs. 4 a, b, 1929.
Variety differing from the typical in the angular form of the
chamber instead of cylindrical.
This variety is very much more abundant than the typical form and
shows considerable variation, from specimens that are but shghtly
angled to those that are very much so. In some specimens there is a
contortion of the chambers, and one is here figured (pl. 9, fig. 7) in
which the chambers have become arranged three in a coil resembling
Flintina. 'The distribution of the variety is given in the following
table:
38 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Taste 9.—Spiroloculina antillarum var. angulata—material examined
Bot-
Num! aba- tom
OM [asl oe Loa Deven |e: | Charest | Secu
HED Se “i ae
Fathoms| ° F.
21940... .-- 1 | H3812 | Entrance to Avatoru Pass, Ra- SIG ee aks wh. co. s. glob. | Rare.
hiroa Atoll, 7.5 miles S. oz. Vol. part.
ZI94T oe 2 | H3814 | Lat. 15° 14” 10” S., long. 147° 51’ OOPS SEE wh. co. s. sh. Do.
5 We glob.
21942___ __. 1 | H3815 | Lat. 15° 15’ S., long. 147° 51’ 35” B24 ho as wh. co. s. brk. Do.
Ww. sh.
2194338 253 1 | H3848 | Village on west side of Niau a2 ieee co. s. glob. oz_--|_ Do.
Atoll, 34 mile E.
21044 22 1 | H3849 | Village on west side of Niau AG 1h een | co. 8S. pter. oz_--| Do.
Atoll, 1.75 miles NE.
21945. .__-. 1 | H3850 | Niau Atoll, 3° S., 14 miles E___- OF tas sees co. Ss. glob. oz_--| Do.
21946* ee 1 | H3855 | Northwest point Apataki, 1 mile 654 | 38.8 | crs. co. S.--..-_.- Do.
SE.
210472 st 2 | H3857 | Center Tikei Island, 44 mile E__ SOO ieee eae dpee <2 Do.
219482 2 =e 6 | H3858 | Ngaruae Pass, Fakarava Atoll, 590) Pemeree eve Re Gore 2 4 1S Few.
28° S., 1 mile E.
21949 2. 1 | H3860 | Southwest end of Fakarava, 2 662 22eee2 co. Ss. pter. oz__-| Rare.
miles NE. |
21950 52 =. 1 | H3905 | Northwest point Hao Atoll, 44 425 | 42 CYSSC0n S222-2 so == Do.
mile SE.
21951 = exes 1 | H3928 |} Anu Anurunga, 1 mile SE__-___- 659 | 38.5 | co. s. brk. sh. Do.
pter. oz.
21952_____- 2 | H3930 | Anu Anuraro Atoll, 34 mile NW_ 438 | 40.7) | COS -22o seo sane Do.
ZIVhs steers 1 |} H3931 | Anu Anuraro Atoll, 4 mile SE_- 405 | 42.5} co. s. pter. oz. Do.
mang. part.
1 Key to abbreviations is given in Table 1.
SPIROLOCULINA ANTILLARUM d’Orbigny AEQUA, new variety
PLATE 9, Fiaures 18 a, bd
Variety differing from the typical in the entirely smooth, polished
surface.
Holotype of variety—Cushman Coll. No. 14739, from the Inside
Lagoon, Pinaki Atoll, Paumotus.
This is the only locality for this variety, but it is fairly common
there.
SPIROLOCULINA ANTILLARUM d’Orbigny var. RETICOSA Chapman
PLATE 9, FiecurE 10
Spiroloculina grata TERQUEM var. reticosa CHAPMAN, Biol. Res. Hndeavour, vol.
3, Di, p: Sls) play igs 254915:
Spiroloculina antillea D’ORBIGNY var. reticosa CUSHMAN, Proc. U. S. Nat. Mus.,
vol. 56, p. 685, 1919.
Variety differing from the typical in the ornamentation of the
surface, which consists of deep elliptical pits, the long axis of the pits
in the long axis of the chambers.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 39
Chapman described this variety from “forty miles South of Cape
Wiles,” and I have had it from material off New Zealand. There
is a single specimen from Albatross Station H38992, Schischmarev
Pass, Marshall Islands, 482 fathoms, which seems to belong to this
variety.
SPIROLOCULINA CADUCA Cushman
PLaTE 9, FrqgurEs 11, 12
Spiroloculina caduca CUSHMAN, Carnegie Inst. Washington Publ. 311, p. 61,
pl. 11, figs. 3, 4, 1922; U. S. Nat. Mus. Bull. 104, pt. 6, p. 42, pl. 9, figs. 1,
2; 1929.
Test broadly elliptical, much compressed, the apertural end ex-
tended; chambers of the adult with a sharp translucent keel, usually
somewhat lobulated; sutures slightly depressed, surface of the
chambers sometimes with irregular, raised costae, more or less oblique
in position; aperture at the end of the cylindrical neck, rounded,
with a simple tooth; surface smooth, shining, but the wall very
thin and brittle. Length, 0.75 mm.; breadth, 0.45 mm.; thickness,
0.1 mm.
The only previous locality for this species is the Tortugas off
Florida. It is interesting to find very typical specimens, two of
which are here figured from Port Lotten, Kersail, Caroline Islands.
SPIROLOCULINA PLANISSIMA (Lamarck) var. SAMOAENSIS Cushman
PLATE 10, Figures 1 a, bd
Spiroloculina planissima (LAMARCK) var. samoaensis CUSHMAN, Carnegie
Inst. Washington Publ. 342, p. 58, pl. 21, figs. 9, 10, 1924.
Variety differing from the typical in the broader form and the
ornamentation consisting of oblique costae running in part way from
the border of the test.
This variety, described and figured from Samoa, occurs in very
typical form at Vavau Anchorage, Tonga Islands. It apparently
is the same as S. striata d’Orbigny, which was named by him in 1826,
but not figured until Fornasini gave figures of those species described
by d’Orbigny but not figured in the 1826 paper. Meanwhile other
authors had used d’Orbigny’s name of striata, so that this is not
available for this form, which is probably somewhat widely dis-
tributed in the Indo-Pacific, d’Orbigny’s species being from Rawack.
SPIROLOCULINA EXIMIA Cushman
PLATE 10, FIGURES 2, 3
Spiroloculina eximia CUSHMAN, Carnegie Inst. Washington Publ. 311, p. 61, pl.
11, fig. 2, 1922; Publ. 342, p. 56, pl. 21, fig. 2, 1924; U. S. Nat. Mus. Bull.
104, pt. 6, p. 42, pl. 8, figs. 7 a, b, 1929.
40 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Test elongate, elliptical, periphery convex, the opposite faces con-
cave, ends of the chamber projecting; sutures fairly distinct, the
outer angle of each chamber projecting above the inner portion of
the adjacent next-formed chamber; wall with a granular, dull sur-
face; apertural end with a projecting cylindrical neck, circular, with
a single, very thin, slightly bifid tooth. Length, 0.5--0.6 mm; breadth,
0.25-0.38 mm; thickness, 0.15-0.18 mm.
This species is not common, but it was found at a number of sta-
tions. It is easily distinguished by the partly rounded periphery of
the chambers and by the projecting lateral margins originally de-
scribed from the Tortugas. It has also been recorded at Samoa and
in the present collections occurs off Levuka, Fiji, 12 fathoms; off
Nairai, Fiji, 12 and 24 fathoms; Viva Anchorage, 3 fathoms; off
Vavau Anchorage, Tonga Islands; and off Rutavu.
SPIROLOCULINA CLARA, new species
PLATE 10, Ficures 4, 5
Test very much compressed, periphery truncate and concave, both
ends somewhat prominently projecting; chambers numerous, narrow,
the sides thickened and opaque, the central portion thin and trans-
lucent; sutures distinct, not much depressed, not limbate; wall
smooth; aperture distinctly exserted, with a tooth or sometimes two
opposite ones. Length, 0.5-0.7 mm.; breadth, 0.3-0.4 mm.; thickness,
0.1 mm.
Holotype.—Cushman, Coll. No. 14781, from Rongelap Atoll, Mar-
shall Islands.
This is probably the same as the form I have figured as “ Spiro-
loculina depressa V’Orbigny.” ®
The species is abundant at the type locality, and is marked by the
peculiar transparency of the middle portion of the chamber giving
TABLE 10.—Spiroloculina clara—material examined
Bot-
Num-
Alba- tom
U.S.N.M. | ber of . Character of Occur-
No. —|speci-| .{TOSs Locally Depth. tenes bottom ! rence
station pera-
i: mens fas
Faihoms | ohne
21935. ..-- 1 | H3875 | Southwest point Tahanae, 269 ene GESuiCOs So ae eo ares
about % mile offshore, 3 miles
NE.
219362. ---- 1 | H8891 | Lat. 16° 30’ S., long. 143° 41’ W_- 540 | 39.7 | co. s. pter.oz_.--| Do.
1Key to abbreviations is given in Table 1.
5U. S. Nat. Mus. Bull. 71, pt. 6, pl. 3, figs. 6 a, b, 1917.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 41
it a peculiar and distinct appearance. It also occurs at Albatross
Stations H38875, one-half mile off southwest point of Tahanae, Pau-
motus, in 269 fathoms; and H3983, one-half mile off entrance to
South Pass, Rongelap, in 400 fathoms. There is a single specimen
that may belong here from Guam Anchorage, Ladrone Islands.
SPIROLOCULINA AFFIXA Terquem
PLATE 10, FicuReEs 6, 7
Spiroloculina afiza TERQUEM, Mém. Soc. Géol. France, ser. 3, vol. 1, p. 55, pl. 5
(10), figs. 13 a-c, 1878.—HowcuHin, Trans. Roy. Soc. South Australia, vol.
12, p. 2, 1889.— CHAPMAN, Journ. Linn. Soc. Zool., vol. 30, p. 16, pl. 1, figs.
23-25, 1907.—CUSHMAN, Proc. U. S. Nat. Mus., vol. 56, p. 685, 1919; U. S.
Nat. Mus. Bull. 100, vol. 4, p. 410, pl. 83, figs. 2 a—c, 1921; Carnegie Inst.
Washington Publ. 342, p. 57, pl. 21, fig. 3, 1924.
Spiroloculina acutimargo H. B. Brapy (in part), Rep. Voy. Challenger, Zool-
ogy, vol. 9, pl. 10, fig. 12, 1884 (not figs. 13-15).—Si1peBorrom (in part),
Journ. Roy. Micr. Soc., 1918, p. 5.
Spiroloculina inaequilateralis ScHLUMBERGER, Mém. Soc. Zool. France, vol. 6,
p. 201, pl. 4, figs. 84-86; fig. 3 (in text), 1893.—SipEBoTtom, Mem. Proc.
Manchester Lit. and Philos. Soc., vol. 54, no. 16, p. 2, pl. 1, fig. 2, 1910.
Test elongate, fusiform, the sides very different from each other,
one side flattened, the other deeply concave; chambers with definite
sutures, which are flush with the surface; wall thin and smooth;
peripheral face somewhat concave, peripheral margin somewhat
carinate, apertural end subacute; aperture elliptical, compressed.
Length, 0.3 mm.; breadth, 0.2 mm.; thickness, 0.1 mm.
There are single specimens, both of which are here figured, from
Albatross Stations H3826, latitude 14° 56’ S., longitude 148° 44’ W.,
in 711 fathoms, off the Paumotus; and H38866, latitude 17° 17’ S.,
longitude 145° 45’ 30’’ W., in 804 fathoms.
TasLe 11.—Spiroloculina affira—material examined
Num) jp fon
U.S.N.M. | ber of 23 ; pao Character of Occur-
No. _ |speci- ee Locality Depth | tem- bottom ! rence
ens pers?
ture
Fathoms| ° F. ‘
210372625 3 1 | H3866 | Lat. 17° 17’ S., long. 145° 45’ 30’ 804 |_.._._| glob. oz. mang_-| Rare,
Ww.
219882 1 | H3829 | Lat. 14° 56’ S., long. 148° 48’ W__| S607 ee wh. co. s. glob. Do.
vol. part.
PAgOoe ae 1 | H3847 | Lat. 16° 08’ S., long. 146° 42’ W__ 609 39 | PMODsO7e-=-s5—s5 Do.
1Key to abbreviations is given in Table 1.
42 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
SPIROLOCULINA sp. (2)
Puate 10, Fiaure 8
There is a single small specimen here figured, rather thick but very
short and nearly circular in outline. It is from Albatross Station
H8876, 1 mile off northwest entrance to Makemo Lagoon, Paumotus.
No other specimens were found to give further details as to the
characters of this form.
Genus HAUERINA d’Orbigny, 1839
Hauerina v’Orzieny, in De la Sagra, Hist. Fis. Pol. Nat. Cuba, “Foraminiféres,”
pp. xxxviii, xxxix, 1839.—H. B. Brapy, Rep. Voy. Challenger, Zoology, vol
9, p. 190, 1884.—CHApMAN, The Foraminifera, p. 97, 1902—-CusHMaN, Cusb-
man Lab. Foram. Res. Spec. Publ. No. 1, p. 150, 1928; U. S. Nat. Mus.
Bull. 104, pt. 6, p. 46, 1929.
Genoholotype—Hauerina compressa d’Orbigny.
Test with the early chambers quinqueloculine, later ones more or
less in one plane, making a half coil, later in some species gradually
shortening so that more than two make up one coil; aperture cribrate.
Tertiary and Recent.
This genus has developed from a quinqueloculine ancestry adding
the later chambers in a single plane and then reducing the length so
that more than two chambers make a coil. With this change in
structure is developed a strongly cribrate aperture.
Some of the species, especially Hauerina CRATES TE, are very com-
mon in the shallow waters of the Indo-Pacific.
HAUERINA FRAGILISSIMA (H. B. Brady)
PLATE 10, FIGURE 9
Spiroloculina fragilissima H. B. Brapy, Rep. Voy. Challenger, Zoology, vol. 9
p. 149, pl. 9, figs. 12-14, 1884.—CuaApman, Journ. Linn. Soc. Zool., vol. 28,
p. 398, 1902.—Daxin, Rep. Pearl Oyster Fisheries Ceylon, p. 230, 1906.
Hauerina fragilissima Mitterr, Journ. Roy. Micr. Soc., p. 610, pl. 18, figs. 8-10,
1898.—HrRon-ALLEN and Eartanp, Trans. Zool. Soc. London, vol. 20,
p. 587, pl. 46, figs. 1, 2, 1915 —CusHman, Proc. U. S. Nat. Mus., vol. 56, p.
638, 1919; Journ. Washington Acad. Sci., vol. 10, p. 200, 1920; U. S. Nat.
Mus. Bull. 100, vol. 4, p. 451, 1921; U. S. Geol. Survey Prof. Paper 129-H, p.
103, pl. 27, fig. 3, 1922; Carnegie Inst. Washington Publ. 342, p. 68, pl. 25,
figs. 2, 3, 1924.
Test much compressed, nearly circular; early chambers quinqueloc-
uline, later ones, which make up the larger part of the adult test,
spiroloculine; periphery rounded; sutures very slightly if at all de-
pressed ; wall very thin, opalescent; the sutures standing out as white
opaque lines; aperture cribrate. Diameter, 0.6 mm.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 43
There are numerous records for this species showing its wide dis-
tribution in the Indo-Pacific region from Kerimba Archipelago off
southeast Africa to the Philippines and southward to Australia.
Very similar specimens occur in the lower Oligocene, Byram marl.
This is easily confused with the flattened forms of Planispirina,
which often occur in the same locality. It may be noted here that
the specimens referred to this species in United States National Mu-
seum Bulletin 71 (pt. 6, pl. 24, fig. 4, 1917) are not Hauerina, but
should be placed in Planispirina.
HAUERINA ORNATISSIMA (Karrer)
PuatTe 10, Ficures 16, 17
Quinqueloculina ornatissima Karrer, Sitz. Akad. Wiss. Wien, vol. 58, p. 151,
pl. 3, fig. 2, 1868.
Hauerina ornatissima H. B. Brapy, Rep. Voy. Challenger, Zoology, vol. 9,
p. 192, pl. 7, figs. 15-22, 1884.—Mutxert, Journ. Roy. Micr. Soc., p. 610, 1898.—
CHAPMAN, Journ. Linn. Soc. Zool., vol. 28, p.178, p. 207 (list), p. 399 (list),
1902.—FornasInI, Mem. Accad. Sci. Istit. Bologna, ser. 5, vol. 10, p. 27,
1902.—Daxin, Rep. Pearl Oyster Fisheries Ceylon, vol. 5, p. 231,
1906.—RuHUMBLER, Zool. Jahrb., Abt. Syst., vol. 24, p. 53, 1906.—Hxrron-
ALLEN and HarLanp, Trans. Zool. Soc. London, vol. 20, pt. 2, p. 590, 1915.—
CusHMAN, U.S. Nat. Mus. Bull. 71, pt. 6, p. 63, pl. 23, figs. 1, 5, 1917; U. S.
Geol. Survey Bull. 676, p. 26, pl. 6, fig. 5, 1918; U. S. Nat. Mus. Bull. 100,
vol. 4, p. 452, 1921; Carnegie Inst. Washington Publ. 311, p. 72, 1922; Publ.
342, p. 67, pl. 24, figs. 7-9, 1924—Hrron-ALLEN and EARLAND, Journ. Linn.
See. Zool., vol. 35, p. 609, 1924—-CusHman, Carnegie Inst. Washington
Publ. 344, p. 82, 1926; U. S. Nat. Mus. Bull. 104, pt. 6, p. 47, pl. 10, figs. 10-
12, 1929.—Hanzawa, Jap. Journ. Geol. Pal., vol. 4, p. 37 (table), 1925
(1926).
Test compressed, early chambers milioline, later ones either em-
bracing and extending in the center to cover the previously
formed chambers or becoming elongate and more or less encircling
the periphery; surface ornamented by strong transverse or radial
ridges or crenulations crossed by numerous fine longitudinal striae;
aperture consisting of numerous small pores in several linear series or
scattered, the sieve plate thus formed extending the full height of the
chamber. Diameter, up to 1.5 mm.
Although this species is typically an Indo-Pacific one, it occurs
sparingly in the West Indian region, but specimens are never so well
developed as in the Pacific and Indian Oceans. Forms referable to
this species also occur in the Oligocene of the Gulf Coastal Plain of
the United States. The types described by Karrer are from the
Miocene of Europe.
In our collections the species is often abundant and occurs at the
following localities: Levuka, Fiji, 12 fathoms; Mokaujar Anchorage,
Fiji; Nairai Islands, Fiji, 12 fathoms; Rotonga, 7 fathoms; Makemo
91513—32——4
44 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Lagoon, Paumotus; Rangiroa ; Port Lotten, Kersail, Caroline Islands;
Rongelap Atoll, Marshall Islands; Guam Anchorage, Ladrone
Islands, 21 fathoms; and the following two Albatross Stations:
H3855, 1 mile off northwest point of Apataki, Paumotus, 654 fath-
oms; and H8967, one-half mile off Maraki Atoll, Marshall Islands,
431 fathoms.
HAUERINA PACIFICA Cushman
PLATE 10, Ficures 10, 11
Hauerina pacifica CusuMAN, U. S. Nat. Mus. Bull. 71, pt. 6, p. 64, pl. 21, fig. 2,
1917.
Test in the early stages quinqueloculine, with strongly angled
chambers, in the adult with the chambers somewhat more inflated
but carinate and irregularly coiled, more than two making up a coil;
sutures fairly distinct in the early portion but not depressed, in the
adult much more depressed; wall smooth, matte; aperture somewhat
projecting with a slightly convex sieve plate, with numerous irregu-
larly arranged pores. Length, 0.7 mm.
The types of this species are from off the Hawaiian Islands in
shallow water ranging from 21 to 33 fathoms where it is common.
In our material specimens occurred at Guam Anchorage, Ladrone
Islands, 21 fathoms, and Mokaujar Anchorage, Fiji Islands.
HAUERINA BRADYI Cushman
PLATE 10, Fieurres 12-15
Quinqueloculina ornatissima Karrer, Sitz. Akad. Wiss. Wien, vol. 58, p. 151,
pl. 3, fig. 2, 1868.—Eaerr, Abh. kén. bay. Akad. Wiss. Miinchen, Cl. II,
vol. 18, p. 244, pl. 3, figs. 9, 10, 23, 24, 1893.—Mitrert, Journ. Roy. Micr.
Soe., 1898, p. 610, pl. 13, fig. 11—CHapmaAn, Journ. Linn. Soc. Zool., vol.
28, p. 899 (list), 1902—RuumeBteER, Zool. Jahrb., Abt. Syst., vol. 24, p. 42,
pl. 3, fig. 39, 1906.—HerrRon-ALLEN and HARLAND, Trans. Zool. Soc. London,
vol. 20, pt. 2, p. 588, 1915; Journ. Linn. Soe. Zool., vol. 35, p. 609, 1924.
Hauerina bradyi CUSHMAN, U. S. Nat. Mus. Bull. 71, pt. 6, p. 62, pl. 23, fig. 2,
1917; Proc. U. S. Nat. Mus., vol. 59, p. 72, 1921; Carnegie Inst. Washington
Publ. 311, p. 71, 1922; Publ. 344, p. 82, 1926; U. S. Nat. Mus. Bull. 104, pt. 6,
p. 47, pl. 10, figs. 4-9, 1929.—CusHMAN and VALENTINE, Contr. Dept. Geol.
Stanford Univ., vol. 1, p. 14, pl. 3, figs. 7 a—c, 1930.
Test much compressed, the very earliest ones milioline, later ones
becoming spiroloculine, and finally, in the last-formed coil, more
than two chambers appear, usually three making up a complete coil;
wall very finely striate-reticulate; periphery rounded or subcarinate;
aperture a sieve plate the entire height of the chamber, curved, with
numerous pores. Diameter, 1 mm.; thickness, 0.15 mm.
This is a widely distributed species in the Indo-Pacific. There is
a very considerable degree of variation shown by the specimens
we have. In the early stages the chambers are in a compact mass of
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 45
a generally quinqueloculine character; following these, chambers are
developed that fail to make the usual half coil, after which, in the
adult, chambers are developed generally in one plane but increasingly
shorter so that three or four make up a complete coil. Some of the
stages in this development are shown in the figures given here.
TABLE 12.—Hauerina bradyi—material examined
| Bot-
Num-
| Alba- tom
U.S.N.M. | ber of . Character of Occur-
No. speci- ees Locality Depth oo bottom 1 rence
mens faire
Fathoms | °F.
22015 ee 1 | H3857 | Center Tikei Island, 44 mile E__ BOO pesos CrS\COn Sass) Rare:
Z201G ee! 1 | H3910 | Southwest point Aki Aki, 1mile E_ 377 4311 COMS=Eee aa eee! Do,
1 Key to abbreviations is given in Table 1.
Genus SIGMOILINA Schlumberger, 1887
Sigmoilina ScHLUMBERGER, Bull. Soc. Zool. France, vol. 12, p. 118, 1887.—
CUSHMAN, Cushman Lab. Foram. Res. Spec. Publ. No. 1, p. 150, 1928;
U. S. Nat. Mus. Bull. 104, pt. 6, p. 48, 1929.
Spiroloculina (part) Costa (not d@’Orbigny), Mem. Accad. Sci. Napoli, vol. 2,
p. 126, 1855 (1857).
Planispirina (part) H. B. Brapy (not Seguenza), Rep. Voy. Challenger, Zool-
ogy, vol. 9, p. 197, 1884.
Genotype.—By designation, Planispirina sigmoidea H. B. Brady.
Test with the early chambers quinqueloculine, later ones added in
planes slightly more than 180° from one another, making a continu-
ously revolving spiral, and in transverse section, producing a sigmoid
appearance; aperture simple with a simple tooth; exterior very often
with a superficial layer of arenaceous material.
Tertiary and Recent.
This genus very evidently developed from Quinqueloculina by the
addition of chambers in planes of more than 180°, so that the result-
ing section is S-shaped.
The following species is the only one of this genus found in our
South Pacific material.
SIGMOILINA EDWARDSI (Schlumberger)
PLATE 11, Ficures 9 a-c
Planispirina (Sigmoilina) edwardsi ScHLUMBERGER, Bull. Soc. Zool. France,
vol, 12, p. 488 (118), pl. 7, figs. 15-18; fig. 8 (in text), 1887.
Sigmoilina edwardsi Hrron-ALLEN and Eartanp, Trans. Zool. Soc. London,
vol. 20, pt. 2, p. 584, pl. 45, figs. 19-21, 1915.—Sipesortom, Journ. Roy. Micr.
Soc., 1918, p. 9 —Hrron-ALLEN and EARLAND, Bull. Soe. Sci. Hist. Nat. Corse,
1922, p. 128; British Antarctic Exped., Zoology, vol. 6, p. 71, 1922; Trans.
Zool. Soe. London, vol. 22, pt. 1, p. 69 (list), 1926.
46 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Test slightly longer than broad, somewhat sigmoid in end view,
the periphery subacute; chambers in the young fairly distinct, in
the highly polished adult somewhat obscured; sutures flush with
the surface, usually indistinct; wall smooth and usually highly
polished; aperture rounded, sometimes with a very slight tooth.
Length, 0.5-0.65 mm.; breadth, 0.4-0.45 mm.; thickness, 0.3-0.35 mm.
This species was found in the Albatross material only from deeper
water and did not occur at any of the shallow stations. The data
for its distribution are given in the accompanying table.
TABLE 18—Sigmoilina edwardsi—méterial examined
Bot-
Num-| alba- tom
Dae caete ross, Locality Depth | tem- ore ot pocrok
mens Deres
ture
fathoms | °F.
219992 --.2 1 | H3819 | Lat. 15° 25’ S., long. 148° 08’ W_-| = 1, 123 |__--_- wh. co. s. glob. | Rare.
vol. part.
S00! a 1 | H3823 | Lat. 15° 01’ S., long. 148° 27’ W-- Sai 2 te. wh. pter.oz.vol.} Do.
part.
DOT 22s ee- 2 | H3824 | Lat. 15° 00’ 20’ S., long. 148° 30’ 850 |____-- wh. glob. oz. Do.
mang. vol.
part.
22002 2S 1 | H3826 | Lat. 14° 56’ S., long. 148° 44’ W_ wl |e wh. pter. oz--.-| Do.
PPOs: - 2ex 1 | H3830 | Lat. 15° 00’ 30’ S., long. 148° 47’ A267 |i wh. co. s. glob. Do.
Ww. vol. part.
22004_ _---- 1 | H3859 | Ngaruae Pass, Fakarava Atoll, 666i 22 2uo8 pter. oz. vol. Do
35° S., 3.5 miles E. part.
DIOOR See 1 | H3866 |} Lat. 17° 17’ S., long. 145° 45’ 30’’ S04}. oes glob. oz. mang--| Do
Ww.
22006 - ----- 1 | H3873 | Southwest point Tahanae, 68° O66 2 See. gues SL Eel bee Do.
N., 4 miles E.
22007 - ----- 5 | H3878 | Lat. 16° 13’ S., long. 148° 48’ W-- O87 ees glob. pter. vol. | Few.
part.
ere é |ssos Tekokoto Atoll, 1 mile E____--- 617))| (38.4 |} Cos SLL =P Rare.
ZOOL e223 1 | H3904 | Lat. 18° 07’ S., long. 141° 26’ W_-| 1, 713 |__---- glob. 'oze. 4 2225 Do.
DOS Lee 1 | H3913 | Northeast end of Nukutavake, 1, 688 | 35.2 | mang. glob_----- Do.
6 miles E.
QUA Bie ae ee, 2 | H3937 | Hereheretue Atoll, 5.3 miles SE_| 1,688 | 35.3 | lt. br. glob. oz. Do.
mang. part.
20142 SE eee 1 | H3977 | Southeast point Elmore Atoll, 1, 283 | 35.9 | ers. glob. oz_---- Do.
30° N., 9 miles W.
1 Key to abbreviations is given in Table 1.
Genus ARTICULINA d’Orbigny, 1826
Articulina p’Orpieny, Ann. Sci. Nat., vol. 7, p. 300, 1826.—H. B. Brapy, Rep.
Voy. Challenger, Zoology, vol. 9, p. 182, 1884——CHapmMan, The Foraminifera,
p. 93, 1902—CusHMAN, Cushman Lab. Foram. Res. Spec. Publ. No. 1, p.
150, 1928; U. S. Nat. Mus. Bull. 104, pt. 6, p. 50, 1929.
Nautilus BarscH (not Linnaeus), Conch. des Seesandes, p. 3, 1791.
Vertebralina (part) Parker, Jones, and H. B. Brapy (not d’Orbigny), Ann.
Mag. Nat. Hist., ser. 3, vol. 16, p. 22, 1865.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 47
Genoholoty pe.—Articulina nitida d’Orbigny.
Test with the early chambers quinqueloculine or triloculine, later
ones in a rectilinear series; aperture in the adult a rounded, usually
elliptical opening, at the end of a short neck with a phialine lip.
Lower Eocene to Recent.
This genus has developed from a quinqueloculine ancestry by the
addition of a uniserial stage, which is represented only by one or two
chambers in some species, but in others these become numerous.
ARTICULINA LINEATA H. B. Brady
PLATE 11, Ficures 1, 2
Articulina lineata H. B. Brapy, Rep. Voy. Challenger, Zoology, vol. 9, p. 183,
pl. 12, figs. 19-21, 1884.—-CusHMaAN, Proc. U. S. Nat. Mus., vol. 59, p. 73, pl.
18, fig. 6, 1921; Carnegie Inst. Washington Publ. 311, p. 70, pl. 12, fig. 4,
1922; U. S. Nat. Mus. Bull. 104, pt. 6, p. 52, pl. 11, figs. 8-10; pl. 12, fig. 1,
1929.
Test much compressed, even in the earlier portion, which is triloc-
uline, the later portion consisting of one or two much compressed
chambers in linear arrangement, periphery usually keeled; chambers
distinct, with numerous fine longitudinal costae; wall translucent,
of a bluish-white color; the aperture with a lip much thickened at
the edge, but not everted and not extending out beyond the periphery
of the chamber. Length, 0.8 mm.; breadth, 0.3-0.4 mm.; thickness,
0.12-0.15 mm.
Brady’s specimens were from off Kandavu, Fiji Islands, 210
fathoms and 255 fathoms, and he also had it from Raine Island,
Torres Strait, 155 fathoms. Brady also recorded the species from
off the Bermudas in 435 fathoms, and according to Nuttall the figured
specimens in the Challenger report are from this last locality.
I have recorded this species from the West Indian region, but a
comparison of the Pacific and the Atlantic series of specimens seems
to show that there are decided differences. If the two are found on
further study to be different, the name /ineata should be applied to
the Atlantic form, as the figured specimens are the Atlantic ones.
In the Pacific collections included in the present paper the most
numerous are from Mokaujar Anchorage, where it is very common.
Similar specimens have occurred also from Levuka, Fiji, in 12
fathoms, but specimens were not obtained from the other area from
which material was examined.
ARTICULINA SULCATA Reuss(7?)
PLate 11, Fieures 3, 4
Recent material from various regions has been referred to the above
species. Specimens consist typically of a quinqueloculine early stage
with one or two linear chambers with a decided neck and the surface
48 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
sharply costate. Such forms are widely distributed and seem never
to have any considerable number of linear chambers. Reuss’s types
were from the Miocene of central Europe and should be compared
with the various forms referred to this species from widely separated
regions. These specimens figured are from Mokaujar Anchorage,
Fiji, and the species occurred also at Albatross Station H3984, off
entrance to South Pass, Rongelap, Marshall Islands, in 746 fathoms.
This is marked as coarse coral sand and undoubtedly represents
material carried out from shallow water.
TABLE 14.—Articulina sulcata—material examined
Bot- |
Num-
U.S.N.M. | ber of| 41P8- ; tom | Character of | Occur-
No. spect Gos Locality, Depth Land bottom rence
ture
Fathoms | ° F.
220175223 1 | H8885 | Southwest point Takume Atoll, BY2) 88.7 CLS) CO, Seto S Rare.
1.5 miles NE.
PAS. 1 | H3890 | Lat. 16° 25’ S., long. 143° 33’ W__| 1,108 | 36.1 | glob. oz. mang__| Do.
1 Key to abbreviations is given in Table 1.
Genus NUBECULINA Cushman, 1924
Nubeculina CUSHMAN, Carnegie Inst. Washington Publ. 342, p. 52, 1924; CusH-
man Lab. Foram. Res. Spec. Publ. No. 1, p. 151, 1928; U. S. Nat. Mus. Bull.
104, pt. 6, p. 55, 1929.
Sagrina (part) H. B. Brapy (not d’Orbigny), Quart. Journ. Micr. Sci., vol. 19,
p. 276, 1879.
Nubecularia (part) H. B. Brapy (not Defrance), Rep. Voy. Challenger, Zoology,
vol. 9, p. 133, 1884.
Genoholotype—Nubecularia divaricata H. B. Brady.
Test elongate, uniserial; initial end coiled or milioline; chambers
distinct, simple; wall imperforate, porcelaneous, with sand grains
attached to the exterior; aperture at the end of an elongated tubular
neck with an everted phialine lip, the apertural opening with a series
of inwardly pointing teeth.
This genus is known only from the Indo-Pacific and is there repre-
sented by several forms. Three of these were found in the material
recorded here. They occur in comparatively shallow water, and some
of the forms reach a large size.
NUBECULINA DIVARICATA (H. B. Brady)
PLATE 11, FicureEs 5, 6
Nubecularia divaricata H. B. Brapy, Rep. Voy. Challenger, Zoology, vol. 9,
p. 136, pl. 76, figs. 11-16, 1884.—A. Sinvestri, Atti Accad. Pont. Nuovi Lincei,
vol. 50, p. 36, 1897.—Mittertr, Journ. Roy. Micr. Soc., p. 261, pl. 5, fig. 4,
1898.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 49
Test composed of subglobular chambers connected by slender cylin-
drical necks; the main body of the chambers with a calcareous base
and the exterior composed of rather coarse calcareous fragments of
various sorts; the apertural end of each chamber with an elongated
tubular neck free from agglutinated material, with a definite lip and
a series of platelike teeth inside the lip.
Brady described this species from the following three stations:
Humboldt Bay, on the north coast of Papua, 37 fathoms; off Raine
Island, Torres Strait, 155 fathoms; and off Tongatabu, Tonga
Islands, 18 fathoms.
Although the species has been referred to by numerous authors,
most of the specimens from the Indo-Pacific belong to the following
variety, which has closely set chambers and a very short neck, and
the teeth in the aperture are quite different. The only specimens we
have that can be referred to Brady’s species are fragments that are
here figured from Nairai, Fiji. They show a characteristic, elongate,
small neck and definite lip.
NUBECULINA DIVARICATA (H. B. Brady) var. ADVENA Cushman
PLATE 11, Fiaures 8 a, b
Nubeculina divaricaia (H. B. Brapy) var. advena CusHMAN, Carnegie Inst.
Washington Publ. 342, p. 53, pl. 19, figs. 1-4, 1924.
Variety differing from the typical in the closely set chambers,
which increase greatly in size and diameter as added; the aperture
tubular, but short and stout with a definite everted phialine lip, and
very coarse teeth in the opening.
Numerous figured specimens such as those of Sidebottom from the
Mediterranean and Heron-Allen and Earland from the Kerimba
Archipelago seem to belong to this variety rather than to the typical
form. I had this variety in abundance from Samoa, where it reaches
a size up to 2.5 mm.
In the present collections it has occurred in typical form from
Mokaujar Anchorage, Fiji; off Levuka, Fiji, 12 fathoms; and off
Nairai, Fiji, 24 fathoms; with somewhat less typical specimens from
off Rotonga (= ? Rarotonga, Cook Islands), 7 fathoms.
NUBECULINA CHAPMANI, new species
PLATE 11, Ficures 7 a, 0
Nubecularia divaricata CHAPMAN (not H. B. Brady), Journ. Linn. Soe. Zool.,
vol. 28, p. 168, pl. 19, fig. 1, 1901.
Chambers somewhat irregularly pyriform, the wall with a cal-
careous base, and the exterior of rather fine fragmentary material;
the apertural end with an elongate, rather stout, cylindrical neck, and
50 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
curved, enlarging toward the base, free from agglutinated material,
the lip very thin, everted, either smooth or with very slight traces of
teeth. Length, 0.8 mm.; diameter, 0.45 mm.
Holotype.—U.S.N.M. No. 22088, from Albatross Station H3898,
one-third mile east of northwest point Hikueru Atoll, Paumotus.
Our figured specimen seems to be almost exactly identical with
that figured by Chapman from the lagoon at Funafuti.
Genus TRILOCULINA d’Orbigny, 1826
Triloculina p’Orpieny, Ann. Sci. Nat., vol. 7, p. 299, 1826—CusHMaAn, Cushman
Lab. Foram. Res. Spec. Publ. No. 1, p. 151, 1928; U. S. Nat. Mus. Bull. 104,
pt. 6, p. 55, 1929.
Miliola (part) LAmMaArcK, Ann. Mus. d’Hist. Nat., vol. 5, p. 351, 1804.
Miliolina (part) Wir1AMmson, Rec. Foram. Great Britain, p. 83, 1858 (and later
authors).
Genotype.—By designation, Miliola trigonula Lamarck.
Test with the early chambers quinqueloculine, at least in the micro-
spheric form, later ones added in the planes 120° from one another,
the third of each series added in the plane of the third preceding and
covering it so that the surface of the test is composed of but three
visible chambers, interior not labyrinthic; aperture simple, typically
with a bifid tooth.
This genus is definitely derived from Quinqueloculina by the addi-
tion of chambers 120° apart and three making up a complete cycle,
and the aperture typically with a bifid tooth. In the microspheric
form all the stages are usually present, but in the megalospheric
form the early quinqueloculine stages may be entirely skipped and
the triloculine stage taken on at once.
In this genus there is a considerable degree of difference in the
characters of the aperture. Some of the species have a large semi-
circular aperture with a flattened tooth, which nearly fills the aper-
ture, while other forms have a narrow aperture with a very narrow
tooth typically bifid at the tip. So far as has been noted these two
forms are not closely related, but they need further study. In the
young stages of the microspheric form chambers are arranged in a
quinqueloculine manner, and such specimens can only be determined
as the young of 7'riloculina by comparison with the adult form that
accompanies them.
TRILOCULINA OBLONGA (Montagu)
PLATE 11, Figures 10 a-c
Vermiculum oblongum Montacu, Test. Brit., p. 522, pl. 14, fig. 9, 1803.
Triloculina oblonga D’OrBiaNy, Ann. Sci. Nat., vol. 7, p. 300, No. 16, 1826;
Modéles, No. 95; in De la Sagra, Hist. Fis. Pol. Nat. Cuba, “ Forami-
niféres,” p. 175, pl. 10, figs. 3-5, 1839.—H. B. Brapy, Trans. Linn. Soe. Zool.,
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS Bk
vol. 24, p. 472 (table), 1864; Nat. Hist. Trans. Northumberland and Dur-
ham, vol. 1, p. 93, 1865 (1867) ; Ann. Mag. Nat. Hist., ser. 4, vol. 6, p. 46,
1870.—TrrRQuEM, Essai Class. Anim. Dunkerque, p. 38, pl. 5, figs. 19 a, b,
1875.—J. WricHt, Proc. Belfast Nat. Field Club, Appendix, p. 103, 1876-
77.—Kiarr, Rep. Norwegian Fish. and Mar. Invest., vol. 1, no. 7, p. 26,
1900.—CuUsHMAN, Contr. Can. Biol., p. 15, 1921 (1922); Carnegie Inst.
Washington Publ. 311, p. 73, 1922; U. S. Nat. Mus. Bull. 104, pt. 6, p. 57, pl.
18, figs. 4, 5, 1929.—CusHmMAN and Moyer, Contr. Cushman Lab. Foram.
Res., vol. 6, p. 52, 1930.—CusHMAN and VALENTINE, Contr. Dept. Geol.
Stanford Univ., vol. 1, No. 1, p. 16, pl. 4, figs. 5, 6, 1930.
Miliolina oblonga Terrier, Atti Accad. Pont. Nuovi Lincei, vol. 33, p. 178,
pl. 1, fig. 2, 1880.—H. B. Brapy, Rep. Voy. Challenger, Zoology, vol. 9,
p. 160, pl. 5, figs. 4 a, b, 1884. (For further references to this species, see
Cushman, U. S. Nat. Mus. Bull. 104, pt. 6, p. 57, 1929.)
Test elongate, the adult with three visible chambers, the last-
formed chamber broadest near the initial end and longer than the
preceding ones; test in end view triangular, the sides broadly curved
and angles rounded; chambers inflated; sutures distinct, depressed ;
wall smooth and usually polished; aperture oval with the tooth
simple or narrow, and bifid at the tip. Length, 0.7-1 mm.; breadth,
0.4-0.55 mm.; thickness, 0.25-0.35 mm.
TaBLE 15.—Triloculina oblonga—material examined
| Bot- |
+ Num-| atpa- m
: ae ao ross. Locality Depth fae cae aoe
mens elas |
ture |
ie sierra ws SN |
Fathoms j° F,
22065 - ---_- 1 | H3809 | Entrance to Avatoru Pass, | G45 | 222 508 {ne. wh. co. s_.--] Rare.
Rahiroa Atoll, 2.5 miles S. | |
22066. - .-.- 1 | H8857 | Center Tikei Island, 34 mile E_- SOOM aoe CYSt COs Ss--seesee Do.
22067 22 | 2 1 | H8876 | Northwest entrance to Makemo A467) ees wh. co. s.------- Do.
Lagoon, 1 mile SE.
22068: 2: 2 1 | H3883 | Northwest Pass Raroia, 5 miles 1,385 | 35.7 | gy. yl. glob. oz. Do.
SE. mang. parts.
220695 = 2 1 | H3884 | Northwest point Raroia, 14 mile 508 | 40.2 | crs. co. s. pter. Do.
SE. OZ.
22070222222 1 | H3898 | Northwest point Hikueru Atoll, 348 | 43.8 | co. s. brk. sh.-._| Do.
14 mile E.
BATE Se se 1 | H3924 |} Nukutipipi Atoll, 1 mile NW--- 649 | 39.0 |_.... dos. eee Do.
220722 2~5- 1 | H8936 | Hereheretue Atoll, 0.3 mile E__- 189 | 62.1 | co.s. mang. part_| Do.
1 Key to abbreviations is given in Table 1.
It is very evident that more than one species is included in the
various references to Montagu’s oblonga. The apertural features
show specimens with an elongate, either simple or bifid tooth, to-
gether with a somewhat similar form with a flattened tooth nearly
filling the aperture. In this South Pacific collection we have lim-
ited this species to those forms which have the narrow tooth.
o2 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Such specimens occur widely distributed and often are very abun-
dant. They occur off Fiji in 40 to 50 fathoms; in 12 to 24 fathoms,
off Nairai; in 3 fathoms, Viva Anchorage; 12 fathoms off Levuka;
and Mokaujar Anchorage. From other areas the species occurs at
Rangiroa; Rutavu; Rotonga; both off the island and from the
lagoon at Niau; Port Lotten, Kersail, Caroline Islands; and in 21
fathoms, Guam Anchorage, Ladrone Islands. The data for the
Albatross stations are given in the accompanying table.
TRILOCULINA CIRCULARIS Bornemann
PuaTE 11, Figures 11 a—c
Triloculina circularis BORNEMANN, Zeitschr. deutsch. geol. Ges., vol. 7, p. 349,
1855.—ReEuvss, Sitz, Akad. Wiss. Wien, vol. 48, pt. 1, p. 41, 1863.—CusHMAN,
U. S. Nat. Mus. Bull. 71, pt. 6, p. 67, pl. 25, fig. 4; pl. 26, fig. 1, 1917; Car-
negie Inst. Washington Publ. 213, p. 290, 1918; Proc. U. S. Nat. Mus., vol.
56, p. 638, 1919; U. S. Nat. Mus. Bull. 100, vol. 4, p. 462, pl. 92, figs. 1, 2, 1921;
Carnegie Inst. Washington Publ. 311, p. 74, 1922.—Horxrr, Flora en Fauna
der Zuiderzee, Protozoa, p. 185, fig. 15 (in text), 1922—CusuMan, Carnegie
Inst. Washington Publ. 342, p. 69, pl. 25, figs. 5, 6, 1924; Publ. 344, p. 82,
1926.—Horker, Zoology of the Faroes, vol. 2, pt. a, Foraminifera, p. 7,
1930.—CUSHMAN and VALENTINE, Contr. Dept. Geol. Stanford Univ., vol. 1,
no. 1, p. 15, pl. 4, figs. 4 a-c, 19380. (For further references to this species,
see Cushman, U. S. Nat. Mus. Bull. 104, pt. 6, p. 58, 1929.)
Test rounded, compressed, the periphery rounded, the three cham-
bers making up the visible portion of the test, rounded, inflated; last-
formed chamber strongly embracing; sutures distinct, depressed;
wall smooth, polished; aperture, a narrow crescentiform slit with a
large, flattened, semicircular tooth. Length, 0.4-1 mm.; breadth,
0.38-1 mm.; thickness, 0.2-0.85 mm.
This is one of the commonest species in warm shallow waters, and
apparently has a very wide distribution. The general character of a
large semicircular platelike tooth nearly filling the large opening is
a constant one. The early quinqueloculine stages are usually found
accompanying the adult triloculine ones, but are usually of much
smaller size.
The accompanying table gives the distribution of this species as
found in our collections at the Albatross stations. In addition it has
occurred in shoal water inside the lagoon, Pinaki Atoll; Vavau
Anchorage, Tonga Islands, 18 fathoms; and off Nairai, Fiji, in 24
fathoms.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 53
TABLE 16.—Triloculina circularis—material examined
Bot-
Num- Alba- tom
ve Stor | apect <tto88 Locality Depth |tem-| Eottomi | ence
mens | St#tion Dare:
Fathoms| ° F.
2 Se 1 | H3813 | Lat. 15° 13’ 10” S., long. 147° 53’ Oa | Soe wh. co. s. glob. | Rare.
10” W. pter.
220508 ae 2% 4 | H3815 | Lat. 15° 15’ S., long. 147° 51’ 35’” B24 y= aes wh. co. s. brk. sh.; Few.
Ww.
S205 ieee | 1 | H3816 | Lat. 15° 16’ 50” S., long. 147° 450 |_----- pter. oz. vol. | Rare.
52’ 30” W. part.
ZA0p 22s 1 | H3848 | Village on west side of Niau 252 ESE ee co. s. glob. 0z..--| Do.
Atoll, 34 mile E.
2205382. 325, 1 | H3855 | Northwest point Apataki, 1 654 | 38.8 | crs, CO. S.-------- Do.
mile SE.
22054... ..- 1 | H3857 | Center Tikei Island, % mile E_- SEOAE (ee 2 dovsi22.225 Do.
20h he Be 3 | H3858 | Ngaruae Pass, Fakarava Atoll, SOOW Reet Re Gon vss Do.
| 28° S., 1 mile E.
22056... =.= | 3 | H3910 | Southwest point Aki Aki, 1 377 | 43 Co; Siisst Sees Do.
| mile E.
22057 - ----- | 4) 3916 | Pinaki Atoll, 1 mile E__-------- 486 | 41 crs. co. s. pter. | Few.
oz.
T2056 eee | 2 | H3931 | Anu Anuraro Atoll, 14 mile SE_- 405 | 42.5 | co. s. pter. oz. | Rare.
| mang. part.
22059 See = 1 | H3935 | Hereheretue Atoll, 1 mile W_--- 594 | 39.5 | ers. co. S_.------- Do.
22060_22..-= 1 | H3967 | Monument, west shore of AST) [3 | be eee Do.
Maraki Atoll, 56° S., 14 mile E.
22061) =. 1 | H3983 | Entrance to South Pass, 400) /|\:43), 4") Cos S82n sess celee: Do.
Rongelap, 4% mile N.
1 Key to abbreviations is given in Table 1.
TRILOCULINA LABIOSA d’Orbigny
PLATE 11, Figures 12 a-c
Triloculina labiosa v’OrBIGNY, in De la Sagra, Hist. Fis. Pol. Nat. Cuba.
“ Foraminiféres,” p. 178, pl. 10, figs. 12-14, 1839—CusHMmAN, U. S. Nat.
Mus. Bull. 71, pt. 6, p. 70, 1917; Proc. U. S. Nat. Mus., vol. 59, p. 70, pl.
16, figs. 13, 14, 1921; Carnegie Inst. Washington Publ. 311, p. 77, pl. 12,
fig. 1, 1922; Publ. 344, p. 83, 1926; U. S. Nat. Mus. Bull. 104, pt. 6, p. 60,
pl. 15, figs. 2, 3, 1929.
Miliolina iabiosa H. B. Brapy, Rep. Voy. Challenger, Zoology, vol. 9, p. 170,
pl. 6, figs. 3-5, 1884.
Test much broader than long, surface largely composed of the
two last-formed chambers; chambers often somewhat irregular;
periphery rounded; surface smooth, but dull; aperture crescentiform,
with a somewhat triangular tooth, placed somewhat back from the
aperture. Diameter, 0.5-0.65 mm.; thickness, 0.2-0.35 mm.
This somewhat irregular species, originally described by d’Or-
bigny from the West Indies occurs in these collections from Ran-
giroa; Rutavu; Hereheretue Island; off Fiji, 40 to 50 fathoms; at
Mokaujar Anchorage, Fiji; and to the northward at Port Lotten,
Kersail, Caroline Islands. At all these stations the material shows
54 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
some variations in the shape of the chambers, which are typically
irregular, but the general characters of the aperture and arrange-
ment of the chambers remain fairly constant.
TRILOCULINA LABIOSA d@’Orbigny SPARSICOSTATA, new variety
PLATE 12, F1acurEs 1 a—c
Variety differing from the typical in the ornamentation of the sur-
face, which has a few, slightly developed, rounding costae with shal.
low depressions between.
Holotype of variety—Cushman Coll. No. 14738, from beach off
wharf, Hereheretue Island.
This variety is abundant at the type locality and occurs as well
with the typical at Rutavu, and a single specimen was taken from
Albatross Station H38924, 7 miles northwest of Nukutipipi Atoll,
Paumotus.
TRILOCULINA IRREGULARIS (d’Orbigny)
PLATE 12, FiauRres 2 a—c
Quinqueloculina irregularis p’ORBIeNy, Ann. Sci. Nat., vol. 7, p. 302, No. 25,
1826.—ForNasini, Mem. Accad. Sci. Istit. Bologna, ser. 6°, vol. 2, p. 67, pl.
3, fig. 14, 1905.
Test slightly longer than broad, nearly circular in side view, com-
pressed, periphery truncate, with the angles produced into definite
costae, apertural end slightly extended; chambers distinct; sutures
distinct, but not depressed; wall matte, with slightly incised short
lines; aperture elongate with a distinct, narrow tooth, with a bifid
tip. Length, 0.8 mm.; breadth, 0.5 mm.; thickness, 0.8 mm.
The types of this species described by d’Orbigny are from the
Mediterranean. Very similar specimens occur in considerable num-
bers in 40 to 50 fathoms off Fiji Islands. In its general characters
it somewhat resembles the one here described as 7riloculina oceanica,
but the characters of the aperture are very distinctive, and the range
of the form with a narrow aperture is very limited.
TRILOCULINA OCEANICA, new species
PLATE 12, Figures 3 a—c
Test about as long as broad, the periphery concave with very
prominent angles to the chambers, which are acute; sutures distinct,
slightly depressed; wall smooth except for the very coarse costae
and angles of the chambers; aperture large, with a semicircular,
flattened tooth nearly filling the opening. Length, 0.5-0.6 mm.;
breadth, 0.45-0.5 mm.; thickness, 0.3-0.35 mm.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 00
Holotype-—U.S.N.M. No. 22078, from Albatross Station H3935,
1 mile west of Hereheretue Atoll, Paumotus.
This species, with its large flat tooth and broad, semicircular open-
ing with the acutely angled chambers, seems to be a distinct one
in fairly deep water among the islands of the South Pacific. There
are specimens from Pinaki Atoll and from the beach off the wharf
at Hereheretue. It occurs also at a number of Albatross stations,
the data for which are given in the accompanying table.
TABLE 17.—Triloculina oceanica—material examined
US.N.M. |berot| trose tom Character of | 0
.S.N.M. |jberof} tross . aracter 0 ecur-
No. |speci-|_ sta- Locality Depth ok bottom ! rence
mens| tion | ture
Fathoms | ° F.
22 o- =~. 22 1 | H3815 | Lat. 15° 15’ S., long. 147° 51’ 35’” BoA ese Se wh.co.s. brk.sh.) Rare.
Wes
|
22074... =~ 1 | H3885 | Southwest point Takume Atoll, BY2) |ioead CrSaC0. Seas ce | Do.
1.5 miles NE.
22075 232= 1 | H3891 | Lat. 16° 30’ S., long. 143° 41’ W__ 540 | 39.7 | co. s. pter.oz_-._-| Do.
22076-..-=-_- 1 | H3905 | Northwest point Hao Atoll, 4% 425 | 42 CTSC0. Se =_-s 8 Do.
mile SE.
220 Tie 1 | H3916 | Pinaki Atoll, 1 mile E--..______- 486 | 41 crs. co. Ss. pter. Do.
Oz.
22078_.----- 1 | H3935 | Hereheretue Atoll, 1 mile W -_-_--} 594°] 39.5 | crs. co. S_---2_-_- Do,
TOMA. 1 | H3936 | Hereheretue Atoll, 0.3 mile E___ 189 | 62.1 | co.s.mang. part_| Do.
1 Key to abbreviations is given in Table 1.
TRILOCULINA OCEANICA, new species, FLINTIANA, new variety
PLATE 12, Ficures 4 a—c
Variety differing from the typical in the ornamentation of the
surface, which consists of numerous additional fine costae, especially
on the periphery, in addition to the angles of the chambers.
Holotype of variety —U.S.N.M. No. 22089, from Albatross Station
H8857, one-half mile east of Center Tikei Island, Paumotus.
This variety is evidently closely allied to the species, but seems
to be worth a distinction from the smoother form.
TRILOCULINA FICHTELIANA d’Orbigny
PLATE 12, FIGURES 6 a-c
Triloculina fichteliana p’OrBieny, in De la Sagra, Hist. Fis. Pol. Nat. Cuba.
“ Foraminiféres,” p. 171, pl. 9, figs. 8-10, 1889.—CusuMan, U. S. Nat. Mus.
Bull “as ptiG; pee OL7 > (Proc, Us 8. Nat: Mus ‘voll’59; ‘p. 70) ple 17;
figs. 1, 2, 1921; Carnegie Inst. Washington Publ. 311, p. 75, 1922; Publ.
344, p. 83, 1926; U. S. Nat. Mus. Bull. 104, pt. 6, p. 63, pl. 17, figs. 1 a—c,
1929.—- CUSHMAN and VALENTINE, Contr. Dept. Geol. Stanford Univ., vol. 1,
no. 1, p. 17, pl. 3, figs. 3 a—c, 1930.
56 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Test subcircular in front view, somewhat compressed, periphery
rounded; chambers distinct; sutures slightly depressed; wall orna-
mented by numerous longitudinal costae; aperture semicircular, with
a slight tooth, which in side view projects beyond the outline of
the aperture, and sometimes shows a distinct perforation. Diameter,
0.55-0.8 mm.; thickness, 0.4-0.45 mm.
This is one of the typical West Indian species described by
d’Orbigny from Cuba and Jamaica. There are numerous records
for it in the general West Indian region, and our material shows
that it extends definitely into the Indo-Pacific, although much of
the material that has been referred to this species is not the same
as the species described by d’Orbigny. The only typical material
we have is from Guam Anchorage, Ladrone Islands, 21 fathoms,
one of which is here figured.
TRILOCULINA SPINATA, new species
PLATE 12, FIGURES 5 a—c
Test elongate, fusiform, periphery rounded, the basal end extend-
ing into a definite pointed spine, the apertural end extended into a
long cylindrical neck; chambers distinct, somewhat inflated; sutures
distinct, not depressed; wall smooth; aperture circular with a very
slight lip and a very small simple tooth, which may or may not be
present. Length, 0.6 mm.; breadth, 0.12 mm.; thickness, 0.1 mm.
Holotype-—Cushman Coll. No. 14633, from Rongelap Atoll, Mar-
shall Islands.
This very distinctive species occurs only at the type locality, but
a number of specimens were present to give the full characters of
the species, which were very constant. The spine at the basal end
is very well developed and is a rather unique feature in this group.
TRILOCULINA TRIGONULA (Lamarck)
PLATE 18, FicuRES 1 a, D
Miliola trigonula LAMARCK, Ann. Mus. d’Hist. Nat., vol. 5, p. 351, No. 8, 1804;
vol. 9, pl. 17, fig. 4, 1807.
Triloculina trigonula, D’ORBIGNYy, Ann. Sci. Nat., vol. 7, p. 299, No. 1, pl. 16,
figs 5-9, 1826; Modéles No. 98, 1826.—CusHMAN, U. S. Nat. Mus. Bull. 104,
pt. 6, p. 56, pl. 12, figs. 10, 11; pl. 13, figs. 1, 2, 1929—CusHMAN and VALEN-
TINE, Contr. Dept. Geol. Stanford Univ., vol. 1, no. 1, p. 16, pl. 4, figs. 7 a-c,
1930.
Miliolina trigonula WILLIAMSON, Rec. Foram. Great Britain, p. 84, pl. 7, figs.
180-182, 1858.—H. B. Brapy, Rep. Voy. Challenger, Zoology, vol. 9, p. 164,
pl. 3, figs. 14-16, 1884.
Test in the adult with three visible chambers, the angles rounded,
the periphery broadly convex, whole test somewhat longer than
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS Oe
wide, in end view rounded, triangular, the sides convex; sutures dis-
tinct; wall smooth; aperture with a rather broad bifid tooth.
Length, 0.5-0.85 mm.; diameter, 0.4-0.45 mm.
As noted previously in Bulletin 104, Lamarck’s type of this spe-
cies is from the Eocene of the Paris Basin, and before the applica-
tion of this name can be very definitely and accurately used it would
be necessary to make sections of microspheric specimens to determine
the characters of the early stages. It has been the usual custom
for authors to place under this species almost any of the triangular
forms that have the edges rounded.
Such specimens as that figured here are fairly common in our
material occurring off Fiji at Mokaujar Anchorage; 12 fathoms off
Nairai; 12 fathoms off Levuka; and also from Makemo beach and
Rutavu, as well as a specimen from Albatross Station H3936, 0.3
mile east of Hereheretue Atoll, Paumotus.
TRILOCULINA AUSTRIACA d’Orbigny
PLATE 138, FIGURES 2 a, 0
Triloculina austriaca DORBIGNY, Foram. Foss. Bass, Tert. Vienne, p. 275, pl. 16,
figs. 25-27, 1846.
Test slightly longer than broad, generally triangular in transverse
section, the sides and periphery of the chambers rounded; chambers
distinct, inflated, usually broader at the base than at the apertural
end, giving an asymmetrical shape to the test in front and side views;
sutures distinct, slightly depressed; wall smooth and polished, but
often marked by very slight elongate pits; aperture either rounded
or somewhat modified as shown in the accompanying figures, with an
elongate tooth bifid at the tip, apertural lip thin and slightly everted.
Length, 0.7 mm.; breadth, 0.45 mm.; thickness, 0.4 mm.
A number of the species described by d’Orbigny from the Miocene
of the Vienna Basin are found still living in the Indo-Pacific region
TABLE 18.—Triloculina austriaca—material examined
Num) ap tout
U.S.N.M. | ber of ae , on Character of Occur-
No. speci- ee Locality Se eee bottom 1 rence
mens Be
ture
Fathoms | ° F.
2AG2- 22225 2 | H8866 | Lat. 17° 17’ S., long. 145° 45’ 30” pV glob. oz. mang_-_| Rare.
Ww.
220830 RE2 es 1 | H3910 | Southwest point Aki Aki, 1 mile 377 AS || CO" Saeco Do.
E.
22064 =.= = 1 |} H3916 | Pinaki Atoll, 1 mile E_-__..____- 486 41 | ers. co. Ss. pter. Do.
Oz. |
|
1 Key to abbreviations is given in Table 1.
58 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
as well as in the Mediterranean. Our material fits very well the
form described by d’Orbigny, which we have from the type locality.
It differs from 7’. trigonula in the side view on account of the greater
inflation of the base, and the chamber is not symmetrical as it is in
T. trigonula. The aperture also is distinctive. Those specimens
that do not have a circular aperture have the sides toward the base
with definite thin lobes, which are difficult to describe but which are
shown in the accompanying figures.
So far as our material shows this species seems to have very definite
and restricted distribution. It is abundant in the collections from
the Paumotu Islands, occurring both inside the lagoon and off the
island of Rongiroa; from the beach off wharf, Hereheretue; on the
outer beach and in the lagoon at Pinaki Island.
Similar specimens occurred at a few Albatross stations, all also off
the Paumotu Islands as shown in the accompanying table.
TRILOCULINA AFFINIS d’Orbigny
PLATE 13, Ficurss 4 a, DB
Triloculina affinis p’OrBIGNY, Ann. Sci. Nat., vol. 7, p. 299, no. 2, 1826.—Trr-
QurEM, Mém. Soc. Géol. France, ser. 3, vol. 2, p. 164, pl. xviii (xv), figs. 1 a-<c,
1882.—ForNASINI, Mem. Acad. Sci. Istit. Bologna, ser. 6*, vol. 2, p. 59, pl. 1,
fig. 1, 1905.
Test slightly longer than broad, triangular in transverse section,
the angles subacute and the outer wall of the chamber decidedly con-
vex; sutures distinct, slightly if at all depressed; wall smooth; aper-
ture generally circular, with a distinctly bifid tooth. Length, 0.35-
0.45 mm.; breadth, 0.35-0.4 mm.
The types of this species were from the Miocene of Dax in the
Bordeaux region. Terquem records a somewhat similar form under
this name, and Fornasini reproduced outline figures from d’Orbig-
ny’s plates. An examination of the Miocene material from the type
locality of Dax shows that there are forms present there that strongly
suggest that this species is closely related to the forms here called
Triloculina trigonula. The outer periphery is distinctly convex, al-
though the peripheral angles are often rather sharply formed.
Such specimens occurred at a number of stations as follows:
Mokaujar Anchorage, Fiji; Viva Anchorage, 3 fathoms, Fiji; Le-
vuka, 12 fathoms, Fiji; near Nairai, Fiji; Rotonga, 7 fathoms;
Makemo Lagoon; and in 21 fathoms off Guam Anchorage, Ladrone
Islands. It occurred also at several Albatross stations as noted in
Table 19.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS
TABLE 19.—Triloculina affinis—material examined
59
Num-| ip tan
U.S.N.M | ber of e ree om | Character of Oceur-
No. speci- Enos Locality Depth | tem- bottom 1! rence
mens Dera
ture
Fathoms | ° F.
22080220... 1 | H3841 | Point Venus, Tahiti Island, 32° Thy eae crs. vol.s.mang.| Rare
S., 4.2 miles W. nod.
2208152 so 1 | H3870 | Village, point Anaa Atoll, 50° 1,110 | 36 fne. co. s. pter. Do.
S., 5 miles W. oz. glob.
220825 = __ =. 1 | H8916 | Pinaki Atoll, 1 mile E__._______- 486 | 41 crs. co. Ss. pter. Do.
oz.
220832525 1 | H3928 | Anu Anurunga, 1 mile SE__-___. 659 | 38.5 | co. s. brk. sh. Do.
pter. oz.
1 Key to abbreviations is given in Table 1.
TRILOCULINA TRICARINATA d’Orbigny
PLATE 18, FiaurEs 3 a, b
Triloculina tricarinata D’ORBIeNY, Ann. Sci. Nat., vol. 7, p. 299, no. 7, 1826;
Modéles, no. 94, 1826.—H. B. Brapy, Trans, Linn. Soc. London, vol. 24, p. 446,
pl. 48, fig. 3, 1864.—CusuMan, U. S. Nat. Mus. Bull. 104, pt. 6, p. 56, pl. 13,
figs. 3 a—c, 1929.
Miliolina tricarinata H. B. Brapy, Rep. Voy. Challenger, Zoology, vol. 9, p. 165,
pl. 3, figs. 17 a, 6b, 1884.
Test in the adult with three visible chambers, the angles of the
chambers sharp, often almost carinate, whole test usually somewhat
longer than wide, in end view triangular, the sides straight; sutures
distinct; wall smooth, often polished; aperture with a narrow bifid
tooth or variously angled in large specimens. Maximum length of
our specimens, 0.6 mm.; diameter, 0.45 mm.
As noted previously, this species, also like affinis, needs definite
sections to determine its early stages in relationships. I have in-
cluded here only those specimens that have very sharp angles and the
sides flat or slightly concave. There is apparently a difference in
the distribution between this and affinis, our specimens being mostly
from Albatross stations in deeper water, although rare specimens
have occurred at Mokaujar Anchorage; Vavau Anchorage, Tonga,
in 18 fathoms; and at Rongelap Atoll, Marshall Islands.
Although there are a great many references to this species, they
are not given here, as so many things have been included under this
name that probably are not strictly to be included under one species.
91513—82. 5
60 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
TABLE 20.—Triloculina tricarinata—material examined
U.S.N.M ean aoe . tom Character of Occur-
“No. — |speci-|_ sta- Ropaitty Bape So bottom 1 rence
mens| tion tare
Fathoms | ° F.
220842228 1 | H3804 | Entrance to Ahii Lagoon, 2.5 15208) azo lt. gy. 0z. glob___| Rare.
miles SE. ;
220854e-. 22 1 | H3818 | Lat. 15° 24’ 10” S., long. 147° 897 jee glob. pter. vol. | « Do.
56’ W. part.
22086_.----- 1 | H3858 | Ngaruae Pass, Fakarava Atoll, 590) cise CSCO $8.28. 2 = Do.
28° S., 1 mile E.
2208 Fe. 2 a 1 | H3873 | Southwest point Tahanae, 68° 966) isk acd glob.oz.mang_-_| Do.
N., 4 miles E.
1 Key to abbreviations is given in Table 1.
TRILOCULINA BERTHELINIANA (H. B. Brady)
PLATH 13, FIGURE 5
Niliolina bertheliniana H. B. Brapy, Rep. Voy. Challenger, Zoology, vol. 9, p.
166, pl. 114, figs. 2 a, b, 1884.—Herron-ALLEN and HARLAND, Trans. Zool.
Soc. London, vol. 20, pt. 17, p. 563, pl. 41, figs. 32-85, 1915; Journ. Linn. Soc.
Zool., vol. 35, p. 605, 1924.
Miliolina tricarinata, reticulated variety, Mittetr, Journ. Roy. Micr. Soc.,
1898, p. 508, pl. 11, fig. 12.
Niliolina tricarinata var. bertheliniana CHAPMAN, Journ. Linn. Soc., Zool., vol.
28, p. 174, 1902.
Triloculina bertheliniana (H. B. Brapy), CUSHMAN, Carnegie Inst. Washington
Publ. 213, p. 290, 1918; U. S. Nat. Mus. Bull. 100, vol. 4, p. 457, 1921.
Test slightly longer than broad, triangular in transverse section;
wall ornamented with numerous, slightly elliptical pits evenly dis-
tributed over the surface of the test, forming a definite pattern. This
is a distinctive Indo-Pacific species so far as the records show. It is
already known from the following localities: Off Ascension Island,
7 fathoms; off Calpentyn, Ceylon, 2 fathoms; shore sands, Tamatavé,
Madagascar; Port Elizabeth, Algoa Bay (H. B. Brady); Malay
Archipelago, rare (Millett) ; Funafuti (Chapman) ; Kerimba Archi-
pelago; Cebu in 45 to 120 fathoms; and Lord Howe Island (Heron-
Allen and Earland); Binang Pool, Subin Bay; Tara Island; Sulu
region off Jolo, off Tawi Tawi, near Basilan, China Sea, off southern
Luzon, and between Burias and Luzon, also Murray Island, Aus-
tralia (Cushman). Our only material of this species, which is very
typical, is from Makemo Lagoon. |
TRILOCULINA BICARINATA d’Orbigny
PLATE 138, FicuREs 6 a—c
Triloculina bicarinata p’ORBIGNY, in De la Sagra, Hist. Fis. Pol. Nat. Cuba,
“ Woraminiféres,” p. 158, pl. 10, figs. 18-20, 1839—CusHMAN, Carnegie
Inst. Washington Publ. 311, p. 76, pl. 12, fig. 7, 1922; Publ. 344, p. 83,
1926; U. S. Nat. Mus. Bull. 104, pt. 6, p. 66, pl. 17, fig. 5, 1929.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 61
Test longer than broad, triloculine, typically with the chambers
with a truncate periphery, the angles somewhat extended; cham-
bers distinct; sutures somewhat depressed; surface ornamented by
reticulations both on the sides and on the outer angles; aperture
elongate, with a definite thin lip, shghtly everted, tooth elongate,
narrow, extending above the outline of the aperture. Length,
0.8-1 mm.; breadth, 0.7-0.9 mm; thickness, 0.55-0.6 mm.
The reticulately ornamented species have been noted under many
different names since the earliest ones given by d’Orbigny. D’Or-
bigny’s Triloculina reticulata was not figured in the 1826 paper, and
while it may apply to various forms with a rounded periphery
such as those to which Brady assigned the name in the Challenger
report, our specimens with definitely angled chambers are much
closer to the West Indian species described by d’Orbigny in 1839
as Triloculina bicarinata.
Our specimens are all from Mokaujar Anchorage, Fiji, where this
form is very abundant, but it was not found in any of the other
stations. There is as usual a considerable degree of variation in
size and arrangement of the reticulations, and in these Fiji speci-
mens the peripheral face of the chambers is highly ornamented,
while the sides are much less so.
TRILOCULINA sp. (7)
PLATE 13, FIGURES 7 a-c
There are in the collections certain forms similar to that here
figured that are very difficult to place specifically. The species seems
to be on the border line between Z7riloculina and Quingueloculina,
and it is figured here in order that attention may be called to this
peculiar form.
Genus PYRGO Defrance, 1824
Pyrgo DEFRANCE, Dict. Sci. Nat., vol. 32, p. 278, 1824—CusHmMan, Cushman
Lab. Foram. Res. Special Publ. No. 1, p. 155, 1928; U. S. Nat. Mus. Bull.
104, pt. 6, p. 67, 1929.
Miliola (part) LAmaArcK, Ann. Mus. d’Hist. Nat., vol. 5, p. 351, 1804.
Biloculina p’Orpiany, Ann. Sci. Nat., vol. 7, p. 297, 1826 (genotype, by desig-
nation, B. bulloides d’Orbigny).
Genoholotype.—Pyrgo laevis Defrance.
In its microspheric form this genus shows all three stages—quin-
queloculine, triloculine, and biloculine—but in the specimens with
a small megalospheric proloculum acceleration takes place and the
quinqueloculine stage is skipped, and in specimens with a very
large proloculum the biloculine character is taken on at once and
both the quinqueloculine and triloculine stages are skipped entirely.
62 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
A number of the species of this genus have become adapted to a
cold, deep-water habitat, an unusual one for most of the genera of
this family.
In the shallow water of the Pacific the genus is often very
abundant, but is limited to a very few species.
PYRGO DENTICULATA (H. B. Brady)
PLATE 14, Ficurns 1-9
Biloculina ringens LAMARCK var. denticulata H. B. Brapy, Rep. Voy. Challenger,
Zoology, vol. 9, p. 148, pl. 3, figs. 4, 5, 1884—Woopwarp, The Observer, vol.
4, p. 76, 1893—MILLeETT, Journ. Roy. Micr. Soc., 1898, p. 262—CHAPMAN,
Journ. Linn. Soc. Zool., vol. 28, p. 398 (list), 1902—Daxin, Rep. Pearl
Oyster Fish. Ceylon, pt. 5, p. 220, 1906.—Hrron-ALLEN and HARLAND, Trans.
Zool. Soc. London, vol. 20, pt. 17, p. 551, pl. 40, figs. 11-18, 1915.
Biloculina denticulata CUSHMAN, U. S. Nat. Mus. Buil. 71, pt. 6, p. 80, pl. 33,
fig. 1, 1917; U. S. Nat. Mus. Bull. 100, vol. 4, p. 476, pl. 98, figs. 3, a, b, 1921;
Carnegie Inst. Washington Publ. 311, p. 78, 1922; Publ. 342, p. 70, 1924;
Publ. 344, p. 88, 1926—Hanzawa, Jap. Journ. Geol. Pal., vol. 4, p. 38,
(table), 1925 (1926).
Pyrgo denticulata CUSHMAN, U. S. Nat. Mus. Bull. 104, pt. 6, p. 69, pl. 18, figs.
3, 4, 1929.
Test elongate, roughly quadrangular in front view, in end view
somewhat compressed, biconvex, apertural end broadly rounded,
opposite end with a series of short, irregular teeth; wall smooth, us-
ually polished; aperture very broad and narrow, extending nearly
the whole width of the test, the ends somewhat expanded, with a long,
narrow tooth, making the inner border of the aperture platelike,
somewhat raised above the level of the surface to which it is attached,
as is the whole border of the aperture. Length, 0.6-1 mm.; breadth,
0.45-0.9 mm.; thickness, 0.3-0.5 mm.
This is a typical species of the Indo-Pacific usually most abundant
in coral reef regions. There are numerous records for it from shal-
low water in various parts of the Pacific from the Philippines to the
Hawaiian Islands, southward through the various groups of islands,
and westward to the coast of Africa. It also occurs in much less
abundance and is less well developed in the Atlantic. It is by far the
most common species of the genus in the material we have examined
from the South Pacific. There is a great deal of variation in this
species, and some of the various forms are figured here. The young
stage is very close to Pyrgo elongata in many ways, and it is these
small light forms that are most easily carried out into deep water by
the current and are found at a number of Albatross stations a short
distance away from these islands, as will be noted. The figures give
these early stages, which may be compared with the adults. Not all
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 63
the specimens show the toothed character of the base, and this
apparently is often lost in the largest, thin-walled specimens. Some
of those with well-developed teeth are here figured as well as the
smooth forms. In these collections the species occurs in typical form
at the following localities: Mokaujar Anchorage, Fiji; Nairai, Fiji,
24 fathoms; Levuka, Fiji, 12 fathoms; Vavau Anchorage, Tonga
Islands, 18 fathoms; Rongelap Atoll, Marshall Islands; Guam An-
chorage, Ladrone Islands, 21 fathoms; Rutavu; and at the following
Albatross Stations: H8857, one-half mile off Center Tikei Islands,
360 fathoms; H3858, Ngaruae Pass, Fakarava Atoll, 599 fathoms;
H3916, 1 mile off Pinaki Atoll, 486 fathoms; H3935, 1 mile off Here-
heretue Atoll, 594 fathoms; H3936, same locality, 189 fathoms; all off
Paumotu Islands; and H3967, one-half mile off Maraki Atoll, Mar-
shall Islands, 431 fathoms,
TABLE 21.—Pyrgo denticulata—material examined
Num-| ip era
, | berof Be : as Character of Occur-
ee NAT speci- SI HEOSS | Locality Depth eo Hottonit meaice
: men: casa
Fathoms | ° F.
22019 Ene 2 | H3857 | Center Tikei Island, % mile E_- 360/28 €TS@ CONSE 2 a2>_ 22] Rare:
22020. -._-- 1 | H8858 | Ngaruae Pass, Fakarava Atoll, SOO OI - ae Gorstacee _ 3 Do.
28° S., 1 mile E.
220 Deters 2 | H3916 | Pinaki Atoll, 1 mile E_..._..__- 486 | 41 ers. co. S. pter. Do.
oz.
22022 = = +2. 1 | H8935 | Hereheretue Atoll, 1 mile W___- 594-|/39. 5 | cers: co. S=/-.=--_]|' Do.
P22 a 1 | H3936 | Hereheretue Atoll, 0.38 mile E___ 189 | 62.1 | co.s.mang. part-} Do.
22024. .._ 1 | H3967 | Monument, west shore of ABT) le ool ae oo oe Do.
Maraki Atoll, 56° S., 44 mile
E., Marshall Islands.
1 Key to abbreviations is given in Table 1.
PYRGO DENTICULATA (H. B. Brady) var. STRIOLATA (H. B. Brady)
PLATE 14, Fieures 10, 11
Biloculina ringens (LAMARCK) var. striolata H. B. Brapy, Rep. Voy. Challenger,
Zoology, vol. 9, p. 148, pl. 3, figs. 7, 8, 1884.——CHAPMAN, Proce. Zool. Soe.
London, p. 8, 1895.—Muttrerr, Journ. Roy. Micr. Soc. p. 262, pl. 5, fig.
8, 1898.—DAkKIN, Rep. Pearl Oyster Fisheries Ceylon, p. 228, 1906——HrEron-
ALLEN and HARLAND, Trans. Zool. Soc. London, vol. 20, p. 551, 1915; Journ.
Linn. Soc. Zool, vol. 35, p. 602, 1924.
Biloculina denticulata (H. B. Brapy) var. striolata CUSHMAN, U. S. Nat. Mus.
Bull. 71, pt. 6, p. 80, pl. 33, figs. 2, 3, 1917; U. S. Nat. Mus. Bull. 100, vol. 4,
p. 477, pl. 98, figs. 2 a—c, 1921; Carnegie Inst. Washington Publ. 311, p. 78,
1922 ; Publ. 342, p. 70, 1924.
Pyrgo denticulata (H. B. Brapy) var. striolata CUSHMAN, U. S. Nat. Mus. Bull.
104, pt. 6, p. 69, pl. 18, figs. 5 a—c, 1929.
64 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Variety differing from the typical in the addition of a surface orna-
mentation consisting of a series of costae on the lower half of the
penultimate chamber, occasionally on the ultimate also.
The only specimens of the variety occur in material from Vavau
Anchorage, Tonga Islands, 18 fathoms; and Mokaujar Anchorage,
Fiji.
PYRGO MURRHINA (Schwager)
PLATE 15, FicurEs 1-3
Biloculina murrhina ScHWAGER, Novara-Expedition, Geology, vol. 2, p. 2083,
pl. 4, figs. 15 a—c, 1866—Munirer-CHALMAS and SCHLUMBERGER, Bull. Soe.
Géol. France, ser. 3, vol. 13, p. 283, figs. 9, 10 (in text) ; p. 290, figs. 15, 16
(in text), 1885.—_ScHLUMBERGER, Mém. Soc. Zool. France, vol. 4, p. 165, pl. 9,
figs. 52, 54; figs. 8, 9 (in text), 1891.—Hecmr, Abh. kén. bay. Akad. Wiss.
Miinchen, Cl. II, vol. 18, pl. 1, figs. 19, 20, 1893——Goés, Bull. Mus. Comp.
Zool., vol. 29, p. 87, 1896.—ScHuBERT, Abh. k. k. Reichs., vol. 20, p. 122,
fig. 17 (in text), 1911.—Prarcey, Trans. Roy. Soc. Edinburgh, vol. 49,
p. 994, 1914—CusHMaAN, U. S. Nat. Mus. Bull. 71, pt. 6, p. 19, fig. 25 (in
text) ; p. 75, pl. 28, fig. 3; pl. 29, fig. 1, 1917; U. S. Nat. Mus. Bull. 100,
vol. 4, p. 470, 1921; Bull. Scripps Inst. Oceanogr., Tech. Ser., vol. 1, p. 140,
1927.
Biloculina depressa D’ORBIGNY var. murrhyna H. B. Brapy, Rep. Voy. Challen-
ger, Zoology, vol. 9, p. 146, pl. 2, figs. 10, 11, 1884.—J. Wriaut, Proc. Roy.
Irish Acad., ser. 3, vol. 1, p. 462, 1891.—CHApman, Proc. Zool. Soc. London,
p. 7, 1895.—Bace, Proce. U. 8. Nat. Mus., vol. 34, p. 117, 1908.—CHarman,
Journ. Linn. Soc. Zool., vol. 30, p. 395, 1910; Zool. Res. Endeavour, pt. 3,
p. 310, 1912; Biol. Res. Endeavour, vol. 3, pt. 1, p. 5, 1915.—SimpEBorrom,
Journ. Roy. Micr. Soc., 1918, p. 9—HrrRon-ALLEN and HARLAND, British
Antarctic Expedition, Zoology, vol. 6, p. 62, 1922.
Pyrgo murrhina CusHMAN, U. 8S. Nat. Mus. Bull. 104, pt. 6, p. 71, 1929.
Test in front view, in young specimens, nearly circular, in adult
specimens somewhat longer than broad, in end view ellipsoidal, with
the borders extended and carinate, the carina interrupted at the
point opposite the aperture, leaving a sinus, rather deep and often
with a long spine at each angle in young specimens; in adults, sinus
less deep, and the spines usually reduced or wanting; wall smooth;
aperture in the young with a neck not exceeding the periphery of
the test; in adults with a prominently exserted tubular neck with a
bifid tooth partially filling the nearly circular opening. Diameter,
up to 1.5 mm.; thickness, 0.5 mm.
This species is widely distributed in deep water in all the oceans
and occurs in the Pliocene of the Indo-Pacific at the type locality,
Kar Nikobar, and in the Pliocene of Fiji.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 65
TABLE 22.—Pyrgo murrhina—material examined
Num-| alba- foe
DARA | oe of tress Locality Depth | tem-| Charsoter of | Ooour
mens Peres
ture
Fathoms | ° F.
220255225 2+ 1 | H3804 | Entrance to Ahii Lagoon, 2.5 1208) je 28 It. gy. oz. glob..-| Rare.
miles SE.
22026254) 1 | H3809 | Entrance to Avatoru Pass, 645 (Se fne. wh.co.s_..-| Do.
Rahiroa Atoll, 2.5 miles S.
TON a 1 | H3820 | Lat. 15° 25’ 59’”’ S., long. 148° 24’ AAR G 2 eo glob. oz. vol. Do.
25° We part.
22028 _ ...-= 1 | H3825 | Lat. 14° 58’ 35’” S., long. 148° 35’ 844, |i. Sue wh. glob. oz. Do.
Ww. mang. vol,
part.
2202094 Foe 1 | H3836 | Lat. 16° 10’ S., long. 148° 26’ W__| 2, 238 |______ vol. m. glob. Do.
mang. nod.
22030__---- 1 | H3841 | Point Venus, Tahiti Island, 32° Won eee crs. vol.s. mang. Do.
S., 4.2 miles W. nod.
77 ise 1 | H3847 | Lat. 16° 08’ S., long. 146° 42’ W_- 609 | 39 glob.0z2s-5---== Do.
2203 2~ = 1 | H3855 | Northwest point Apataki, 1 mile 65451038" 8) |fersxcouss_ 2.----| Do:
SE.
29033225 -= 1 | H3878 | Lat. 16° 13’ S., long. 143° 48’ W_- 987 | ie bute glob. pter. vol. Do.
part.
22034-—.-.. 1 | H3903 | Lat. 18° 08’ S., long. 141° 49’ W__ 2,187 | 35.2 | vol. m. glob_---. Do.
99035- ====- 1 | H3996 | Tekokoto Atoll, 1 mile E_._____. OLY G8. 40 COL S250 2s) 223 Do.
22036_____- 1 | 113922 | Lat. 20° 31’ S., long. 142° W____- Dae ah [aac edie eee] | Do.
Dai cee 1 | H3923 | Nukutipipi Atoll, 5 miles NW-_-| 2,315 | 35 rdcces glob PDOs
990382 oo a= 1 | H3991 | Lat. 9° 40’ S., long. 169° 32’ W.__| 1, 583 | 35.5) |p GlOpwiOZs saoaeen= Do.
1 Key to abbreviations is given in Table 1.
PYRGO GLOBULA (Bornemann)
PLATE 15, F1icuRES 6-8
Biloculina globulus BorNEMANN, Zeitschr. deutsch. geol. Gesell., vol. 7, p. 349,
pl. 19, fig. 8, 1855.—ScHLUMBERGER, Mém. Soc. Zool. France, vol. 4, p. 188,
pl. 12, figs. 97-100; figs. 42-44 (in text), 1891—-CHapmMaAN, Subantarctic
Islands of New Zealand, p. 317, 1909.—CusHMAN, U. S. Nat. Mus. Bull. 71,
pt. 6, p. 78, pl. 31, fig. 2, 1917; Bull. 100, vol. 4, p. 474, pl. 95, figs. 2 a, b, 1921.
Test in front view oval, in end view irregularly globular; chambers
very tumid; suture slightly depressed, in side view the last-formed
chamber at the aboral end somewhat curving out over the preceding
chamber; wall smooth, white; aperture semicircular, with a simple
flat tooth filling a large part of the opening. Length, about 1 mm.
Specimens that seem to be the same as the specimens described by
Bornemann from the Oligocene of Europe, and recorded by Schlum-
berger from off the Azores, are found at numerous stations in the
Albatross collections but not in the shallow-water samples. The
typical form has a flat tooth largely filling the aperture. The speci-
mens figured from the Philippines do not have this typical tooth but
are otherwise similar.
66 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
TABLE 23.—Pyrgo globula—material examined
Num! atba- en
oN. {spect| TOSS Locality Depth |tem-| Bottom” | ‘tenos
mens peas
ture
Fathoms| ° F.
22039... ... 1 | H3815 | Lat. 15° 15’ S., long. 147° 51’ 35’” B24 spas wh. co. s. brk. | Rare.
W. sh.
22040. =... 1 | H3825 | Lat. 14° 58’ 35’’ S., long. 148° 35’ 844i eels wh. glob. oz. Do.
AW mang. vol.
part.
22041 Lae 1 | H3828 | Lat. 14° 51’ 20’ S., long. 148° 51’ 624, ELTA Wh? CO.S2220e-5 Do.
20’ W.
220424 ei 5u 1 | H3841 | Point Venus, Tahiti Island, 32° C75: | 22a crs. vol.s. Mang. Do.
S., 4.2 miles W. nod.
22043 eee 1 | H3853 | Pakaka entrance to Apataki 613" |"30.44\rcO avOlZ= 23-222 Do.
Lagoon, 50° N., 2 miles E.
20044 1 | H3857 | Center Tikei Island, 44 mile E__- SEOn/ 2.2684 CTSHCONS. hae scan Do.
22045 == 1 | H3860 | Southwest end of Fakarava, 2 GODS Mest co. Ss. pter.0z----| Do.
miles NE.
22046 422-2 1 | H3876 | Northwest entrance to Makemo AGT Gs baw whitcors 425... Do.
Lagoon, 1 mile SE.
22047... =. 1 | H3914 | Northeast point Nukutavake, 1 636) "S8s9p| tease ee ss oe Do.
mile S.
22048... 1 | H3954 | West end of Nomuka Island, 33° 600 | 39.2 | co.s. pum. pter. Do.
N., 6 miles E. oz.
1 Key to abbreviations is given in Table 1.
PYRGO MILLETTII (Cushman)
PLATE 15, Fiaures 4, 5
Miliolina durandii Mirtetrr (part), Journ. Roy. Micr. Soc., 1898, p. 268, pl. 6,
figs. 8-10 (not fig. 7) —HERoN-ALLEN and HaRrLanp, Trans. Zool. Soc. Lon-
don, vol. 20, p. 565, pl. 42, figs. 11-16, 1915.
Biloculina millettii CusHMan, U. S. Nat. Mus. Bull. 71, pt. 6, p. 81, pl. 34,
figs. 4, 5, 1917; Carnegie Inst. Washington, Publ. 311, p. 77, 1922; Publ. 342,
p. 71, 1924.
Pyrgo millettii CUSHMAN, U. S. Nat. Mus. Bull. 104, pt. 6, p. 68, pl. 19, fig. 1,
1929.
Test in front view, broadly elliptical, in end view, compressed ;
chambers biconvex; the periphery with a definitely developed carina;
wall smooth, except for occasional transverse ribs usually indistinct ;
aperture slightly produced, broadly elliptical, with a slightly thick-
ened border joining the carina at its outer edge; the aperture with
a small bifid tooth.
This is a typical Indo-Pacific species of wide distribution and ap-
parently extends to the West Indies region, where it is known only
from the Tortugas. In the Pacific it is recorded as follows: Malay
Archipelago (Millett); Kerimba Archipelago, southeast Africa,
Burma, Queensland, Java, Macassar, and Tahiti (Heron-Allen and
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 67
Earland) ; Hongkong Harbor, Philippines, and Samoa (Cushman).
The only specimens found in all the samples examined in the present
collections are from Levuka, Fiji, in 12 fathoms, but these are very
typical.
Family OPHTHALMIDIIDAE
Test calcareous, imperforate, early chambers at least planispiral,
except in degenerate forms; wall without an arenaceous coating;
aperture typically open, without a tooth.
Subfamily CORNUSPIRINAE
Test made up of a proloculum and an elongate, planispiral, tubular
second chamber.
Genus CORNUSPIRA Schultze, 1854
Cornuspira ScHULTzZE, Organismus Polythal., p. 40, 1854.—H. B. Brapy, Rep.
Voy. Challenger, Zoology, vol. 9, p. 198, 1884.—CHaApMAN, The Foraminifera,
p. 99, 1902.—CusHMAN, Cushman Lab. Foram. Res. Spec. Publ. No. 1, p. 160,
1928; U. S. Nat. Mus. Bull. 104, pt. 6, p. 78, 1929.
Orbis (part) PHitiep1, Enum. Moll. Siciliae, vol. 2, p. 147, 1844.
Operculina (part) CzszeK, Haidinger’s Nat. Abh., vol. 2, p. 146, 1848.
Spirillina (part) WiLL1Amson, Rec. Foram. Great Britain, p. 91, 1858.
Genotype.—By designation, Cornuspira planorbis Schultze.
Test consisting of a proloculum followed by a long planispirally
coiled second chamber, rounded or complanate; wall calcareous, im-
perforate; aperture formed by the open end of the chamber, some-
times constricted and with a thickened lip.
This genus is represented in the South Pacific by but a single
species in our collections.
CORNUSPIRA INVOLVENS Reuss
PLATE 16, FIGURES 2 a, B
For complete references to this species see Cushman, U. S. Nat. Mus. Bull. 104,
pt. 6, pp. 80-81, 1929.
Test nearly circular in side view, consisting of a proloculum and a
long closely coiled, planispiral second chamber of nearly equal diame-
ter throughout, slightly involute; suture distinct, somewhat de-
pressed; wall smooth and polished, occasionally showing slight lines
of growth; aperture nearly the size of the open end of the tube.
Diameter of our South Pacific specimen, not exceeding 0.35 mm.
This is a common and widely distributed species, but in our mate-
rial never reaches any considerable size. Specimens are common off
Fiji near Nairai, 12 and 24 fathoms; off Levuka, 12 fathoms; off
68 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Vavau Anchorage, 3 fathoms; and Mokaujar Anchorage. Speci-
mens are also common off Niau and in Niau Lagoon; Vavau Anchor-
age, Tonga Islands, 18 fathoms; Rotonga, 7 fathoms; Rongelap
Atoll, Marshall Islands; Port Lotten, Kersail, Caroline Islands; and
Guam Anchorage, Ladrone Islands, 21 fathoms.
Subfamily NODOBACULARIINAE
Genus NODOBACULARIA Rhumbler, 1895
Nodobacularia RHUMBLER, Nachr. Ges. Wiss. Gottingen, 1895, p. 87.—CusHMAN,
Cushman Lab. Foram. Res. Spee. Publ. No. 1, p. 164, 1928; U. S. Nat. Mus.
Bull. 104, pt. 6, p. 86, 1929.
Nubecularia (part) Jones and PaRgKer (not Defrance), Quart. Journ. Geol. Soc.,
vol. 16, p. 455, 1860.
Genoholotype—Nubecularia tibta Jones and Parker.
Test with a globular proloculum followed by a planispiral tubular
second chamber, the adult chambers in a rectilinear series; aperture
simple, with a lip.
Lias to Recent.
This genus was apparently developed early in the history of the
family, and species and specimens are common in the Jurassic.
There are few living species.
The early stages in most specimens are very delicate, and easily
broken away so that the majority of specimens have only the later
heavy-walled chambers present. In some specimens there is only the
proloculum and a second coiled chamber before the rectilinear series
is taken on. In others there are two or three coiled chambers usually
making up a half coil or less, but the proloculum is always evident.
This is a typically tropical shallow-water genus. Owing to the early
chambers being usually lacking, it is difficult to place some of the
early described species, such as, for example, the forms usually re-
ferred to Articulina conico-articulata. The original figure given by
Batsch does not have the early chambers, and it is very difficult to be
sure of his species.
NODOBACULARIA ANTILLARUM Cushman PACIFICA, new variety
PLATE 16, FiagurEs 1 a—c
Test slender, elongate, often slightly curved; the early chambers
gradually increasing in length as added, somewhat pyriform, slightly
wider toward the base; sutures distinct, much depressed; wall orna-
mented by numerous, usually 13 to 15, sharp, raised, longitudinal
cost; apertural end slightly contracted, and then expanded into a
very definite lip, which is smooth. Length, 2.2 mm.; breadth, 0.2 mm.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 69
Holotype of variety—Cushman Coll. No. 14741, from Mokaujar
Anchorage, Fiji.
This variety has also occurred off Levuka, Fiji, in 12 fathoms, and
off Nairai, Fiji, 12 fathoms. Somewhat less typical specimens, but
probably belonging to this same variety, occurred off Rotonga, 7
fathoms, and at Vavau Anchorage, Tonga Islands, 18 fathoms.
It is close to the species I have described as Nodobacularia antil-
larwm from the West Indian region. The Pacific specimens, how-
ever, uniformly have more and sharper costae, and the earliest cham-
bers are more compressed and smaller in comparison with the adult
chambers.
NODOBACULARIA MILLETTI, new species
Pratp 16, Fiaures 3, 4
Articulina conico-articulata Mitterr (not Batsch), Journ. Roy. Micr. Soc.,
1898, p. 511, pl. 12, figs. 9, 10.
Test with the early chambers planispiral, the later ones rectilinear
and rapidly increasing in size as added, adult chambers very strongly
pyriform, greatest breadth at the base, and thence rather rapidly
contracting toward the apertural end, which is expanded into a
broad smooth lip; sutures distinct; wall ornamented by a few, 12
to 18 usually, high, thin, platelike costae, which terminate in spines
at the basal end, those of the later chambers often bifurcating;
aperture elliptical, with a very broad lip. Length, 1.5 mm.; breadth,
0.5 mm.; thickness, 0.35 mm.
Holotype——Cushman Coll. No, 14743, from Mokaujar Anchorage,
Fiji.
This is probably the same form as that described and figured by
Millett in the above reference from the Malay Archipelago. His
specimens, however, show a much larger proloculum, and the coiled
chambers show practically no compression, while in our speci-
mens from Fiji there is a strongly compressed young stage. Later
chambers, however, seem to be very close to ours. It may be that
his specimens are the megalospheric form, while ours are micro-
spheric. This is a fine distinctive species, beautifully ornamented,
and so far as the material we have shows, has only occurred off Fiji
at Mokaujar Anchorage, and off Levuka in 12 fathoms. It is
common at the type locality.
In the early stages there may be developed short chambers of one-
third of a coil in length, which begin to take on the characteristic
costz ornamentation before the adult uniserial stage is developed.
70 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Subfamily OPHTHALMIDIINAE
Genus SPIROPTHALMIDIUM Cushman, 1927
Spiropthalmidium CusHMAN, Contr. Cushman Lab. Foram. Res., vol. 3, p. 37,
1827; Cushman Lab. Foram. Res. Spec. Publ. No. 1, p. 165, 1928; U. S.
Nat. Mus. Bull. 104, pt. 6, p. 90, 1929.
Spircloculina (part) H. B. Brapy (not d’Orbigny), Rep. Voy. Challenger, Zool-
ogy, vol. 9, p. 154, 1884.
Genoholotype.—S piroloculina acutimargo H. B. Brady (part).
Test similar to Ophihalmidium, but accelerated, the stage having
two chambers in a coil quickly reached; plate between the chambers
usually present; aperture simple, without teeth.
There is a single species of this genus occurring in our material.
SPIROPTHALMIDIUM ACUTIMARGO (H. B. Brady)
PLATE 16, FIGURES 5 a, b
Spiroloculina acutimargo H. B. Brapy (part), Rep. Voy. Challenger, Zoology,
vol. 9, p. 154, pl. 10, fig. 18 (mot figs. 12, 14, 15), 1884——Batkwitt and
J. Wricut, Trans. Roy. Irish Acad., vol. 28, Sci., p. 328, 1885.—Sippa.,
Proc. Lit. Phil. Soc. Liverpool, 1886, p. 72 (list) —Hatkyarp, Trans. Man-
chester Micr. Soc., 1889, p. 59—Cuastrr, lst Rep. Southport Soc. Nat. Sci.,
1890-91, p. 55, 1892.— J. Wricut, Proc. Roy. Irish Acad., ser. 3, vol. 1, p.
463, 1891.—Mittert, Journ. Roy. Micr. Soc., 1898, p. 264—-CHAPMAN, Journ.
Linn. Soe. Zool., vol. 28, p. 172, 1901.—SipesottomM, Mem. Proc. Manchester
Lit. Philos. Soc., vol. 48, no. 5, p. 6, 1904—Har.Lanp, Journ. Quekett Micr.
Club, ser. 2, vol. 9, p. 192, 1905.—Baae, Proc. U. S. Nat. Mus., vol. 34,
p. 119, 1908—CHapman, Journ. Linn. Soc. Zool., vol. 30, p. 396, 1910.—
Heron-ALLEN and HarLanp, Proc. Roy. Irish Acad., vol. 31, pt. 64, p. 24,
pl. 1, fig. 8, 1918.—CHapman, Biol. Res. Endeavour, vol. 3, pt. 1, p. 6, 1915.—
Herron-ALLEN and HARLAND, Trans. Zool. Soc. London, vol. 20, pt. 17, p. 557,
1915; Trans. Linn. Soe. London, ser. 2, vol. 11, p. 208, 1916.—CusHMAN, U.S.
Nat. Mus. Bull. 71, pt. 6, p. 31, pl. 5, fig. 1, 1917—Sipesorrom, Journ. Roy.
Micr. Soc., 1918, p. 5—Cusuman, U. S. Nat. Mus. Bull. 100, vol. 4, p. 398,
1921; Carnegie Inst. Washington Publ. 342, p. 56, 1924.
Spiropthalmidium acutimargo CusHMAN, Contr. Cushman Lab. Foram. Res.,
vol. 3, p. 37, pl. 8, fig. 5, 1927; Cushman Lab. Foram. Res. Special Publ.
No. 1, pl. 20, fig. 7; pl. 21, fig. 5, 1928; U. S. Nat. Mus. Bull. 104, pt. 6,
p. 90, pl. 22, fig. 1, 1929.
Test oval, much compressed, planispiral throughout, peripheral
margin carinate; development consisting of a proloculum followed
by several coils of an undivided second tubular chamber, then by
chambers a half coil in length separated by a wide flange; wall
smooth; aperture rounded, without a tooth. Length, 0.5 mm.;
breadth, 0.3 mm.; thickness, 0.05 mm.
The only specimens of this species are from three Albatross sta-
tions as given in Table 24. All the specimens are very typical.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS ts
TABLE 24,.—Spiropthalmidium acutimargo—material examined
Num-| aip tot
U.S.N.M. | ber of Be . om | Character of Occur-
No. —|speci-| .{f0SS Locality Depth | tem- bottom rence
mens station pera-
ture
Fathoms | ° F.
2209025225 2 | H3829 | Lat. 14° 56’ S., long. 148° 48’ W_- $60), -Be2s" wh. co. s. glob. | Rare.
vol. part.
22001 - == 1 | H3830 | Lat. 15° 00’ 30” S., long. 148° 47’ 2b falseaeee ans doA 5s: Fo Do.
Ww.
22092 === 1 | H3873 | Southwest point of Tahanae, S6Gn|2--= 22 glob. oz. mang__ Do.
68° N., 4 miles E.
Genus PLANISPIRINA Seguenza, 1880
Planispirinad Seevuenza, Atti R. Accad. Lincei, ser. 3, vol. 6, p. 310, 1880.—H. B.
Brapy, Rep. Voy. Challenger, Zoology, vol. 9, p. 193 (in part), 1884.—
CHAPMAN, The Foraminifera, p. 98, 1902.—CusHMAN, Cushman Lab. Foram.
Res. Spec. Publ. No. 1, p. 166, 1928; U. S. Nat. Mus. Bull. 104, pt. 6, p. 93,
1929.
Genotype.—By designation, Planispirina communis Seguenza.
Test in the early stages like Cornuspira, later divided into cham-
bers, several to a coil; aperture simple, without a tooth.
There are two species of this genus in our Pacific material, only
one of which is at all common.
PLANISPIRINA EXIGUA (H. B. Brady)
PLATE 16, FiauRres 7 a, b
Hauerina exigua H. B. Brapy, Quart. Journ. Micr. Sci., vol. 19, p. 53, 1879.
Planispirina exigua H. B. Brapy, Rep. Voy. Challenger, Zoology, vol. 9, p. 196,
pl. 12, figs. 14; text-fig. 5b, 1884—H. B. Brapy, Parker, and Jonzgs, Trans.
Zool. Soc. London, vol. 12, p. 216, pl. 40, fig. 4, 1888—Howcutin, Trans.
Proc. Roy. Soc. South Australia, vol. 12, p. 5, 1889.—Eacrr, Abh. kin. bay.
Akad. Wiss. Miinchen, Cl. II, vol. 18, p. 245, pl. 3, figs. 11, 12, 1893—
Woopwarp, The Observer, vol. 4, p. 77, 1893——Mutuerr, Journ. Roy. Mier.
Soc., p. 611, pl. 13, fig. 18, 1898 —CuApman, Journ. Linn. Soe. Zool., vol. 30,
p. 21, pl. 2, fig. 43, 1907; Subantarctic Islands of New Zealand, p. 323, 1909.—
HERON-ALLEN and HarLanp, Trans. Zool. Soc. London, vol. 20, p. 590, 1915.—
CHAPMAN, Biol. Res. Endeavour, vol. 3, pt. 1, p. 11, 1915—Cusuman, U. S.
Nat. Mus. Bull. 71, pt. 6, p. 9, fig. 7 (in text), 1917—Hrron-ALLEN and
EARLAND, Journ. Roy. Micr. Soe., p. 1383, 1924—CusuHMan, U. S. Nat. Mus.
Bull. 104, pt. 6, p. 94, 1929.
Test much compressed, planispiral, the early whorls undivided,
later ones showing usually two or three chambers to the coil, rather
indistinct except when the specimens are wet; sutures rather indis-
tinct except those of the last-formed coil; wall fairly smooth, usually
opaque; aperture a narrow elongate opening without a definite tooth.
Diameter, 0.3 mm.; thickness, 0.03-0.05 mm.
72 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
This is a small easily overlooked species occurring in its typical
form rather widely distributed in the Indo-Pacific, although there
are records from other regions. I have in the present collections
very typical specimens from 12 fathoms off Levuka, Fiji, where it
is fairly common, and from near Nairai, Fiji. A single specimen
also occurred at Albatross Station H3812, 714 miles south of entrance,
Avatoru Pass, Rahiroa Atoll, Paumotus.
PLANISPIRINA AURICULATA Egger
PLATE 16, FIGURES 6 a—c
Planispirina auriculata Eacrr, Abh. kén. bay. Akad. Wiss. Mtinchen, Cl. II, vol.
18, p. 245, pl. 3, figs. 18-15, 1898 —-Hmron-ALLEN and HARLAND, Trans. Zool.
Soe. London, vol. 20, p. 590, pl. 46, figs. 3-7, 1915.—SipeBottom, Journ. Roy.
Micr. Soc., 1918, p. 10.—CusHMAN, Carnegie Inst. Washington Publ. 311,
p. 62, pl. 10, fig. 8, 1922—Hrron-ALLEN and HaArLanp, Bull. Soc. Sci. Hist.
Nat. Corse, 1922, p. 123; Journ. Linn. Soc. Zool., vol. 35, p. 609, 1924.—
CusHMAN, U.S. Nat. Mus. Bull. 104, pt. 6, p. 93, pl. 22, fig. 3, 1929.—CusH-
MAN and Parker, Proc. U. 8. Nat. Mus., vol. 80, art. 3, p. 6, pl. 2, fig. 3, 1931.
Test minute, generally oval in outline, biconvex, consisting of a
few milioline chambers, the apertural end somewhat extended, turned
so that the aperture is entirely at one side of the test; aperture
elliptical or rounded, with a broad, flaring lip; sutures fairly dis-
tinct, not depressed; wall translucent; color, bluish white. Length,
0.3 mm.; breadth, 0.15 mm.; thickness, 0.05 mm.
The only specimens we have are a few from Port Lotten, Kersail,
Caroline Islands,
This minute species is, like the preceding, very easily overlooked on
account of its small size. From the records, however, it has a wide
distribution in both the Atlantic and Pacific. It is easily distin-
guished by the large aperture in proportion to the size of the test,
and the fiaring lip.
Genus VERTEBRALINA d’Orbigny, 1826
Vertebralina p’OrBiIeNy, Ann. Sci. Nat., vol. 7, p. 283, 1826-——H. B. Brapy, Rep.
Voy. Challenger, Zoology, vol. 9, p. 1884.—CHAPpMAN, The Foraminifera,
p. 97, 1902.—CusHMAN, Cushman Lab. Foram. Res. Spec. Publ. No. 1, p.
168, 1928; U. S. Nat. Mus. Bull. 104, pt. 6, p. 95, 1929.
Genoholotype—Veriebralina striata d’Orbigny.
Test with the early chambers planispiral, later ones in a rectilinear
series; aperture simple, a long narrow slit either at the outer end of
the chamber or somewhat laterally placed, typically with a definite
lip. Eocene to Recent.
In the present collection the genus is represented only by the fol-
lowing species, strata.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 73
VERTEBRALINA STRIATA d’Orbigny
PLATE 16, Fieures 8-10
Vertebralina striata p’OrgiaNy, Ann. Sci. Nat., vol. 7, p. 288, No. 1, Modéles,
No. 81, 1826.—WiLL1aAMson, Rec. Foram. Great Britain, p. 90, pl. 7, figs.
196a, b(=197, 198), 1858.—W. B. CARPENTER, PARKER, and Jones, Introd.
Foram., p. 72, pl. 5, figs. 17-25, 1862_—Parker, Jones, and H. B. Brapy, Ann.
Mag. Nat. Hist., ser. 3, vol. 16, p. 32, pl. 1, fig. 1, 1865; ser. 4, vol. 8, p. 239,
pl. 8, fig. 27, 1871.—Scuwacer, Boll. R. Com. Geol. Ital., vol. 8, p. 27, pl., fig.
106, 1877.—H. B. Brapy, Rep. Voy. Challenger, Zoology, vol. 9, p. 187, pl. 12,
figs. 14-16, 1884——A. Sitvesrri, Atti Accad. Sci. Acireale, vol. 7, p. 22,
1896.—Muuerr, Journ. Roy. Micr. Soc., p. 607, pl. 138, fig. 1, 1898—Smpr-
BoTToM, Mem. Proc. Manchester Lit. Philos. Soc., vol. 48, no. 5, p. 18, 1904.—
Daxin, Rep. Pearl Oyster Fish. Ceylon, p. 231, 1906.—CHApMAN, Journ.
Quekett Micr. Club, ser. 2, vol. 10, p. 125, 1907.—Sipesorrom, Mem. Proc.
Manchester Lit. Philos. Soc., vol. 54, no. 16, p. 6, 1910.—HrRoN-ALLEN
and BARLAND, Journ. Roy. Micr. Soc., p. 305, 1911; Trans. Zool. Soc. Lon-
don, vol. 20, p. 587, 1915—CusuMaAn, U. S. Nat. Mus. Bull. 71, pt. 6, p.
38, pl. 22, figs. 3, 4, 1917—Marrtinorri, Atti Soc. Ital. Sci. Nat., vol. 59,
p. 327, fig. 170 (in text), 1920—CusuMan, U. S. Nat. Mus. Bull. 100, vol.
4, p. 414, 1921—HerEron-ALLEN and Earuanp, Bull. Soc. Sci. Hist. Nat.
Corse, 1922, p. 123.—CusHMAN, Carnegie Inst. Washington Publ. 342, p.
58, 1924; U. S. Nat. Mus. Bull. 104, pt. 6, p. 96, pl. 22, figs. 6a, b, 1929.
Test much compressed, early portion close coiled, later uncoiling
in a series of low, broad, uniserial chambers; sutures distinct, de-
pressed; surface ornamented by fine longitudinal striae; aperture
elongate, irregular, the lip on one side being shorter than on the
other, making the aperture really on the ventral side of the test, lip
smooth, ends rounded and not projecting. Length, 0.6-1.2 mm.;
breadth, 0.5-0.7 mm.; thickness, 0.2-0.25 mm.
This is a common species in the material, rather widely distributed,
and while it varies somewhat in form, because the early stages are
close coiled and the later ones somewhat uncoiled, nevertheless
the ornamentation, which is very constant, will serve to distinguish
the species from any other of those allied to it. There are occa-
sional specimens that tend toward V. reticulosa Cushman described
from the Philippines, but none of them has these characters very
strongly developed. D’Orbigny’s model of this species shows a much
uncoiled specimen with the sides in the adult nearly parallel. Such
specimens are not common, but are occasionally found in large series
of specimens. Most of those found are usually in the early coiled
stages. There are specimens in the present collection from Mokaujar
Anchorage, Fiji; Levuka, Fiji, 12 fathoms; Viva Anchorage, Fiji,
3 fathoms; and off Nairai, Fiji, 12 fathoms. Specimens also occur
off Vavau Anchorage, Tonga Islands; off Rotonga (= ?Rarotonga) ;
and Guam Anchorage, Ladrone Islands, in 21 fathoms.
74 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Subfamily NUBECULARIINAE
Genus PARRINA Cushman, 1931
Parrina CusHMAN, Contr. Cushman Lab. Foram. Res., vol. 7, p. 20, 1931.
Silvestria ScuuBert, Pal. Zeitschr., vol. 3, p. 166, 1920—CusHmMan, Cushman
Lab. Foram. Res. Spec. Publ. No. 1, p. 169, 1928.
Nubecularia (part) of authors.
Genoholotype—Nubecularia inflata H. B. Brady (not Terquem) =
NV. bradyi Millett.
Test with the early chambers where visible irregularly coiled, later
chambers inflated and very irregular, sometimes coiled, sometimes
in an irregular linear arrangement; wall calcareous, imperforate;
aperture variable in shape, often rounded, and irregularly placed.
Recent.
This genus is represented in the Indo-Pacific in shallow water of
coral reef regions by the following species, brady.
PARRINA BRADYI (Millett)
PLATE 17, FIcurEs 1—4
Nubecularia inflata H. B. BrRapy (not Terquem), Rep. Voy. Challenger, Zoology,
vol. 9, p. 135, pl. 1, figs. 5-8, 1884.—A. Sitvestri, Atti Accad. Sci. Acireale,
vol. 7, p. 30, 1895-96; Mem. Accad. Pont. Nuovi Lincei, vol. 50, p. 35, 1897.—
Baae, Proc. U. S. Nat. Mus., vol. 34, p. 116, 1908—CusHMAN, U. S. Nat.
Mus. Bull. 104, pt. 6, p. 98, 1929.
Nubecularia bradyi Mitterr, Journ. Roy. Micr. Soc., 1898, p. 261, pl. 5, figs. 6a,
b.—CHAPMAN, Journ. Linn. Soe. Zool., vol. 28, p. 169, pl. 19, fig. 3, 1900.—
SrpmEBoTToM, Mem. Proc. Manchester Lit. Philos. Soe., vol. 48, no. 5, p. 3,
1904.—RuHuUMBLER, Zool. Jahrb., Abt. Syst., vol. 24, p. 40, pl. 2, figs. 17-19,
1906.—CHAPMAN, Journ. Quekett Micr. Club, ser. 2, vol. 10, p. 119, 1907.—
SipesotTom, Mem. Proc. Manchester Lit. Philos. Soc., vol. 54, no. 16, p. 2,
1910.—HeERON-ALLEN and HARLAND, Journ, Roy. Micr. Soe., 1911, p. 300.—
Prarcrey, Trans. Roy. Soc. Edinburgh, vol. 49, p. 993, 1914-——-CHAPMAN,
Victorian Nat., vol. 32, p. 49, fig. 1 (in text), 1915—Hrron-ALLHN and
HARLAND, Trans. Zool. Soc. London, vol. 20, p. 550, pl. 40, figs. 8-10, 1915.—
CUSHMAN, U. S. Nat. Mus. Bull. 71, pt. 6, p. 41, pl. 8, figs. 4, 5, 1917;
Proc. U. 8. Nat. Mus., vol. 56, p. 636, 1919.—HrRon-ALLEN and EARLAND,
British Antarctic Expedition, Zoology, vol. 6, p. 60, 1922; Journ. Linn.
Soe. Zool., vol. 35, p. 601, 1924; Trans. Zool. Soc. London, vol. 22, pt. 1,
p. 68 (list), 1926.
Silvestria bradyi ScuHuspertT, Pal. Zeitschr., vol. 3, p. 166, 1920.—CusHMAN,
Cushman Lab. Foram. Res. Spec. Publ. No. 1, pl. 58, figs. 11-13, 1928.
Parrina bradyi CusHMAN, Contr. Cushman Lab. Foram. Res., vol. 7, p. 20,
1931.
Test very irregular in form, the early portion often showing signs
of coiling but early becoming irregular in form and shape; chambers
inflated, globular, irregularly arranged, either in a loose coil or in a
more or less linear series; wall smooth, calcareous, imperforate,
polished ; aperture rounded, or very irregular in form and irregularly
placed. Diameter of test, up to 1 mm.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS wo
This species seems to be widely distributed in warm coral reef
areas, especially in the Mediterranean and the Indo-Pacific. It is
an extremely variable form, and assumes many shapes.
Our specimens are all from comparatively shallow waters, except
for one station off Fiji in 21 fathoms or less. They include the fol-
lowing stations: Off Nairai, Fiji, 12 fathoms; off Levuka, Fiji, 12
fathoms; off Fiji, 40-50 fathoms; Rangiroa; off Niau; Makemo
Lagoon; beach off wharf, Hereheretue Island; Rutavu; Vavau An-
chorage, Tonga Islands, 18 fathoms; Port Lotten, Kersail, Caroline
Islands; and Guam Anchorage, Ladrone Islands, in 21 fathoms.
Family FISCHERINIDAE
Test coiled, earlier ones somewhat planispiral, later ones trochoid,
all coils visible from the dorsal side, only the last-formed one from
the ventral side; chambers distinct but not inflated, usually four or
five making up the last-formed coil; wall calcareous, imperforate;
aperture rounded, formed by the open end of the last-formed
chamber.
This family is represented by the single genus Fischerina, which
is commonest in the warm waters of the Tropics. The family is
closely related to and derived from the family Ophthalmidiidae.
It is a direct derivative of the genus Cornuspira, assumes a trochoid
form, and is divided into chambers.
Genus FISCHERINA Terquem, 1878
Fischerina TerquEM, Mém. Soc. Géol. France, ser. 3, vol. 1, p. 80, 1878.—CusH-
MAN, Cushman Lab. Foram. Res. Spec. Publ. No. 1, p. 170, 1928; U. S.
Nat. Mus. Bull. 104, pt. 6, p. 100, 1929.
Genoholotype.—Fischerina rhodiensis Terquem.
Test coiled in a low conical spiral; chambers few in each coil, all
visible from the dorsal side, only those of the last-formed coil from
the ventral side; wall calcareous, imperforate; aperture formed by
the open end of the last-formed chamber, simple.
The species of this genus are mostly confined to shallow warm
water of the Tropics.
Our material has yielded three species of this genus, all of them
apparently different from the West Indian one.
FISCHERINA PELLUCIDA Millett
PLATE 17, FIGURES 7 a—c
Fischerina pellucida MittetTt, Journ. Roy. Micr. Soc., 1898, p. 611, pl. 18, figs. 14,
15.—HERON-ALLEN and EARLAND, Trans. Zool. Soe. London, vol. 20,
p. 591, 1915—CusuHMAN, Proc. U. S. Nat. Mus., vol. 56, p. 636, 1919; Carnegie
Inst. Washington Publ. 342, p. 53, pl. 19, fig. 9, 1924—Hrron-ALLEN and
HARLAND, Journ. Linn. Soe. Zool., vol. 35, p. 610, 1924.
915183—32——-6
76 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Test forming a very low, compressed, trochoid spiral of only about
114 to 2 whorls, the earliest portion in the microspheric form with
the chamber following the proloculum, making about one complete
coil, followed by an adult whorl, in which there are from 5 to 6 cham-
bers on the dorsal side, the coils very slightly if at all overlapping, on
the ventral side the central portion slightly umbilicate, the last-
formed whorl making up most of the surface on the ventral side,
periphery broadly rounded; wall thin, translucent, bluish white;
aperture circular, formed by the open end of the last-formed chamber.
Diameter, 0.25 mm.; thickness, 0.05-0.07 mm.
Millett’s types of this species are from the Malay Archipelago, and
there are records for it, references to which are given above, from
the Kerimba Archipelago (Heron-Allen and Earland) ; off New Zea-
land, off Samoa (Cushman) ; Lord Howe Island (Heron-Allen and
Earland). Rather typical specimens occurred in our material from
Port Lotten, Kersail, Caroline Islands, and from Guam Anchorage,
Ladrone Islands, in 21 fathoms.
FISCHERINA HELIX Heron-Allen and Earland
PLATE 17, Ficures 5 a-c
Fischerina helic Hrron-ALLEN and HARLAND, Trans. Zool. Soc. London, vol.
20, pt. 17, p. 591, pl. 46, figs. 10-14, 1915; British Antarctic Expedition,
Zoology, vol. 6, p. 73, 1922.
Test forming a definite trochoid spiral of 4 or 5 volutions, pe-
riphery broadly rounded, whorls with about 3 or 4 chambers to each;
sutures flush with the surface except the spiral suture, which is
somewhat depressed; wall very thin, fragile, opalescent; aperture
formed by the open end of the last-formed chamber. Diameter,
0.25-0.8 mm.; height, 0.18-0.2 mm.
Heron-Allen and Earland described this species from the Kerimba
Archipelago off southeast Africa, and also recorded it from a
dredging off the Poor Knights Islands, New Zealand. The only
material I have had in these present collections is 40 to 50 fathoms
off Fiji.
FISCHERINA INVOLUTA, new species
PuatEy 17, Figures 6 a-c
Test nearly planispiral and almost completely involute on both
sides; chambers somewhat indistinct, three or four in the final
whorl; sutures sigmoid on the ventral side, on the dorsal side radial,
flush with the surface; wall very thin, fragile; aperture semicircular,
formed by the open end of the last-formed chamber. Diameter,
0.3-0.35 mm.; thickness, 0.2 mm.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 77
Holotype —uvU.S.N.M. No. 14626, from Rangiroa.
This species differs from the other known ones of the genus in
having the whorls almost completely involute, especially in the later
development. A few specimens seem to show that the earliest cham-
bers are more open. The ventral side seems to be almost completely
involute with a very slight umbilical depression, the dorsal side
flattened or slightly convex. As far as our material shows, this is
the commonest species of the genus in the South Pacific. It is
abundant at the type locality, and also occurs at Rutavu; Makemo
Lagoon, Paumotus; inside lagoon, Pinaki Atoll; beach off wharf,
Hereheretue; and from Albatross Station H3933, latitude 20° 02’
S., longitude 144° 28’ W., Paumotus.
Family TROCHAMMINIDAE
Subfamily TROCHAMMININAE
Genus TROCHAMMINA Parker and Jones, 1860
TROCHAMMINA TURBINATA (H. B. Brady)
PLATE 17, Ficurns 8 a, b
Haplophragmium turbinatum H. B. Brapy, Quart. Journ. Micr. Sci., vol. 21,
p. 50, 1881; Rep. Voy. Challenger, Zoology, vol. 9, p. 312, pl. 35, figs. 9a—c,
4884.
Trochammina turbinatum Himer and Ficxert, Zeitschr. Wiss. Zool., vol. 65, p. 695,
1889.—CusHMAN, U. S. Nat. Mus. Bull. 104, pt. 2, p. 81, pl. 17, fig. 2, 1920;
Bull. Seripps Inst. Oceanogr., Tech. Ser., vol. 1, no. 10, p. 142, 1927.—
CUSHMAN and Moyer, Contr. Cushman Lab. Foram. Res., vol. 6, p. 53, pl. 7,
fig. 10, 1930.
Test spiral, early portion regular, low spired, last-formed whorl in
the adult often becoming oblique; chambers 5 to 8 in the last whorl,
umbilicate; wall arenaceous, usually smoothly finished, cement yellow-
ish brown; aperture a narrow slit at the base of the ventral side of
the chamber. Diameter, 0.35 mm.; thickness, 0.18 mm.
The only specimens of this species are from Albatross Station
H3973, latitude 5° 20’ S., longitude 169° 43’ W., in 2,411 fathoms.
Subfamily GLOBOTEXTULARIINAE
Genus NOURIA Heron-Allen and Earland, 1914
Nouria Heron-ALLEN and EArtanp, Trans. Zool. Soe. London, vol. 20, pt. 12,
p. 375, 1914—CusHMAN, Smithsonian Mise. Coll., vol. 77, no. 4, p. 30, 1925;
Cushman Lab. Foram. Res. Spec. Publ. no. 1, p. 174, 1928.
Genotype—By designation, Nouria polymorphinoides Heron-
Allen and Earland.
78 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Test free, of several chambers, irregularly arranged; wall arena-
ceous; aperture simple, terminal.
Recent.
There is a single species present in the collections.
NOURIA POLYMORPHINOIDES Heron-Allen and Earland
PLATE 17, Fiaures 9 a, 0
Nouria polymorphinoides HrRon-ALLEN and HAriAnp, Trans. Zool. Soc. London,
vol. 20, pt. 12, p. 376, pl. 37, figs. 1-14, 1914; pt. 17, p. 615, 1915 —CusHMan,
Proc. U. 8S. Nat. Mus., vol. 56, p. 601, pl. 75, figs. 4, 5, 1919.—HERoN-ALLEN
and EARLAND, British Antarctic Expedition, Zoology, vol. 6, p. 103, 1922.—
CusHMAN, Contr. Cushman Lab. Foram. Res., vol. 3, p. 189, 1927; Bull.
Seripps Inst. Oceanogr., Tech. Ser., vol. 1, p. 142, 1927.
This species has now been recorded from several widely distributed
stations in the Indo-Pacific region from the east coast of Africa,
the western coast of America, and from off New Zealand. The
record given by Heron-Allen and Earland from the Eocene of Biar-
ritz® seems from the figures to be a different thing. The Indo-
Pacific material is definite in its general characters. Our specimens
are from Mokaujar Anchorage, Fiji, in comparatively shallow water.
They often have sponge spicules incorporated in the test as well as
other fragments of various sorts. These are usually neatly cemented,
and there are traces of sutures in most of the specimens.
6 Halkyard, Mem. Proc. Manchester Lit. Philos. Soc., vol. 62, pt. 2, pl. 1, figs. 6, 7, 1918
(1919).
Fieures 1-38.
4a, b.
5a, 0:
6-10.
11, 12
Figures 1-3.
4a, b.
Bad
8-10.
Fieures 1, 3:
2, 4.
5 a, b.
6 a, 6.
€ a,b.
10, 11.
EXPLANATION OF PLATES
PuaTe 1
Reophaz agglutinatus Cushman. X70. Fig. 1, Young specimen.
Albatross H3910. Fig. 2. Albatross H3919. Fig. 3, Specimen
broken, showing interior, Albatross H3779.
Haplophragmoides trullissata (H. B. Brady). X 70. a, Side
view; b, peripheral view. Albatross H3779.
Spiroplectammina milletti (Cushman). X 40. a, Front view;
b, apertural view. Albatross H3959.
Textularia foliacea Heron-Allen and Earland. X 40. Fig. 6,
Vavau Anchorage, Tonga Islands. Fig. 7, a, Front view; 5, side
view. Levuka, Fiji, Fig. 8, a, Front view; b, apertural view.
Mokaujar Anchorage, Fiji. Fig. 9, a, Front view; b, side view.
Rongelap Atoll, Marshall Islands. Fig. 10, Mokaujar Anchor-
age, Fiji.
Textularia foliacea Heron-Allen and Earland oceanica, new
variety. Fig. 11, Holotype, X 30. a, Front view; 6, side
view. Mokaujar Anchorage, Fiji. Fig. 12, Young stage.
x 40. Rotonga.
PLATE 2
Textularia semialata Cushman. X 40. a, Front view; b, aper-
tural view. Mokaujar Anchorage, Fiji.
Textularia candeiana d’Orbigny. X 40. a, Front view; }, aper-
tural view. Makemo Lagoon, Paumotu Islands.
Textularia agglutinans d’Orbigny. X 55. Fig. 5, Rongelap
Atoll, Marshall Islands. Fig. 6, a, Front view; b, apertural
view. Levuka, Fiji. Fig. 7, Albatross H3961.
Textularia conica d’Orbigny. X 55. a, a, a, Front view; b, b,
b, apertural views. Figs. 8, 10, Mokaujar Anchorage, Fiji.
Fig. 9, Vavau Anchorage, Tonga Islands.
PLaTE 3
Textularia conica d’Orbigny. X 55. a, Front view; b, apertural
view. Rongelap Atoll, Marshall Islands.
Textularia corrugata Heron-Allen and Earland. X 40. a, Front
view; b, apertural view. Off Fiji, 40-50 fathoms.
Textularia albatrossi Cushman. X 40. a, Front view; b, aper-
tural view. Albatross H3858.
Textularia concava Karrer. X 55. a, Front view; b, apertural
view. Albatross H3815.
Gaudryina triangularis Cushman var. angulata Cushman. X55.
a, Front view; b, apertural view. Mokaujar Anchorage, Fiji.
. Gaudryina bradyi Cushman. X 55. a, a, Front views; b, b, aper-
tural views. Fig. 8, Albatross H3818. Fig. 9, Albatross H3901.
Gaudryina quadrangularis Bagg. X 40. a, a, Front views; b, b,
apertural views. Albatross H3959.
79
80 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
PLATE 4
Fiaures 1a, b. Gaudryina rugulosa, new species. X 25. a, Front view; 6,
apertural view. Alofi Niue.
2 a, b. Gaudryina sp. (?). X 55. a, Front view; 6b, apertural view.
Makemo Lagoon, Paumotu Islands.
3 a, b. Clavulina communis d’Orbigny, X 25. a, Front view; b, apertural
view. Albatross H3816.
4a, b. Clavulina pacifica Cushman. X 30. a, Front view; b, aper-
tural view. Makemo Lagoon, Paumotu Islands.
5, 6. Clavulina difformis H. B. Brady. X 25. a, a, Front views;
b, b, apertural views. Fig. 5, Guam Anchorage, Ladrone
Islands. Fig. 6, Mokaujar Anchorage, Fiji.
7 a, b. Clavulina pacifica Cushman. X 30. a, Front view; }, aper-
tural view. Rangiroa.
8 a, b. Clavulina difformis H. B. Brady. X 25. a, Front view; 3,
apertural view. Vavau Anchorage, Tonga Islands.
9a, b. Clavulina pacifica Cushman. X 30. a, Front view; 6, aper-
tural view. Rangiroa.
10 a, b. Clavulina difformis H. B. Brady. a, Front view; 6, aper-
tural view. Guam Anchorage, Ladrone Islands.
PuateE 5
Fiaures 1 a-c. Quinqueloculina anguina Terquem var. agglutinans (Wiesner).
xX 55. a, b, Opposite sides; c, apertural view. From off Nai-
rai, Fiji.
2-4. Quinqueloculina berthelotiana d’Orbigny. X 55. Figs. 2, 4,
From Rangiroa. Fig. 3, a, 6, Opposite sides; c, apertural
view. From Hereheretue Island.
5. Quinqueloculina bidentata d’Orbigny. XX 40. Off Levuka, Fiji.
6, 7. Quinqueloculina costata d’Orbigny. X 55. a, a, b, b, Opposite
sides; c,c, apertural views. Vavau Anchorage, Tonga Islands.
8 a-c. Quinqueloculina crassa d’Orbigny var. subcuneata Cushman.
X 55. a, 6, Opposite sides; c, apertural view. Rongelap
Atoll, Marshall Islands.
9,10. Quinqueloculina funafutiensis (Chapman). X 70. Fig. 9, Mo-
kaujar Anchorage, Fiji. Fig. 10, a, 6, Opposite sides; c, ap-
ertural view. Off Niau, Paumotu Islands.
11 a-c. Quinqueloculina crenulata, new species. XX 55. a, b, Opposite
sides; c, apertural view. From Port Lotten, Kersail, Caro-
line Islands.
12. Quinqueloculina cf. gracilis d’Orbigny. > 40. Rangiroa.
PLATE 6
Fieures 1 a-c. Quinqueloculina gualtieriana d’Orbigny. X 55. a, b, Opposite
sides; c, apertural view. Vavau Anchorage, Tonga Islands.
2 a-c. Quinqueloculina lamarckiana d’Orbigny. X 40. a, b, Opposite
sides; c, apertural view. Off Levuka, Fiji.
3, 4. Quinqueloculina parkeri (H. B. Brady). X 30. Fig. 3, a, b,
Opposite sides. From Guam Anchorage, Ladrone Islands.
5, 6. Quinqueloculina polygona d’Orbigny, X 40. a, a, b, b, Opposite
sides; c, c, apertural views. Fig. 5, Rotonga. Fig. 6, Off
Levuka, Fiji. ,
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS $1
Figures 1 a-c.
2 a, b.
34,
5-8.
FIGURE i
2 a, b.
3 a, b.
4a, b.
5 a, b.
627.
8 a, b.
9-11.
FiauREs 1 a, b.
2 a, b.
3-5.
6-9.
10.
i eb
13 a, b.
PuatTe 7
Quinqueloculina samoaensis Cushman. X 40. a, 6, Opposite
sides; c, apertural view. Off Levuka, Fiji.
Quinqueloculina semireticulosa, new species. X 55. a, Front
view; b, apertural view. Port Lotten, Kersail, Caroline
Islands.
Quinqueloculina striatula, new species. X 40. Fig. 3, Holotype.
a, b, Opposite sides; c, apertural view. Mokaujar Anchorage,
Fiji.
Quinqueloculina sulcata d’Orbigny. XX 40. Figs. 5, 8, Off Fiji,
40—50 fathoms. 8a, 6, Opposite sides; c, apertural view. Figs.
6, 7, Mokaujar Anchorage, Fiji. 7a, b, Opposite sides; c, aper-
tural view.
PLATE 8
Schlumbergerina alveoliniformis (H. B. Brady). XX 25. Mo-
kaujar Anchorage, Fiji.
Massilina australis, new species. X 40. a, Front view; }, aper-
tural view. Rotonga.
Massilina alveoliniformis Millett. X< 40. a, Front view; },
apertural view. Off Levuka, Fiji.
Massilina macilenta (H. B. Brady). X 55. a, Front view; },
apertural view. Off Levuka, Fiji.
Massilina crenata (Karrer). X 55. a, Front view; b, apertural
view. Albatross H3858.
Massilina inaequalis Cushman. X 40. Fig. 6, From Viva An-
chorage, Fiji. Fig. 7, a, Front view; b, apertural view. Vavau
Anchorage, Tonga Islands.
Massilina planata, new species. X 25. a, Front view; },
apertural view. From Guam Anchorage, Ladrone Islands.
Spiroloculina grateloupi d’Orbigny. X 40. Fig. 9, a, Front view;
b, apertural view. Mokaujar Anchorage, Fiji. Fig. 10, Viva
Anchorage, Fiji. Fig. 11, Albatross H3930.
PuaTE 9
Spiroloculina gratelowpi d’Orbigny serrulata, new variety. X
55. a, Front view; b, apertural view. Viva Anchorage, Fiji.
Spiroloculina grateloupi d’Orbigny acescata, new variety. X
40. a, Front view; b, apertural view. Off Nairai, Fiji.
Spiroloculina antillarum d’Orbigny. X 50. Fig. 3, a, Front
view; b, apertural view. Rangiroa. Fig. 4, Albatross H3853.
Fig. 5, Albatross H3910.
Spiroloculina antillarum d’Orbigny var. angulata Cushman.
<x 50. Fig. 6, a, Front view; 6, apertural view. Guam An-
chorage, Ladrone Islands. Fig. 7, Deformed, triserial form.
Niau Lagoon, Paumatus. Figs. 8, 9, Off Fiji, 40-50 fathoms.
Spiroloculina antillarum d’Orbigny var. reticosa Chapman.
xX 60. Albatross H3992.
Spiroloculina cauduca Cushman. X 50. Fig. 12, a, Front view;
b, apertural view. Port Lotten, Kersail, Caroline Islands.
Spiroloculina antillarum d’Orbigny aequa, new variety. X
50, a, Front view; b, apertural view. Inside Lagoon, Pinaki
Atoll, Paumotu Islands.
82 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Figures 1 a, b.
2,3.
4, 5.
Gyize
16, 17.
Ficures_ 1, 2.
3, 4.
5, 6.
ees:
8 a, b.
9 a-c.
10 a-c.
11 a-c.
12 a-c.
Ficures 1 a-c.
2 a-c.
Puate 10
Spiroloculina planissima (Lamarck) var. samoaensis Cushman.
X50. a, Front view; b, apertural view. Vavau Anchorage,
Tonga Islands.
Spiroloculina eximia Cushman. XX 55. Fig. 2, a, Front view;
b, apertural view. Levuka, Fiji. Fig. 3, Viva Anchorage,
Fiji.
Spiroloculina clara, new species. X 55. Fig. 4, Holotype. a,
Front view; b, apertural view. Rongelap Atoll, Marshall
Islands.
Spiroloculina afixa Terquem. X 75. Fig. 6, a, b, Opposite
sides; c, apertural view. Albatross H3826. Fig. 7, Alba-
tross H3866.
. Spiroloculina sp. (?). X 75. Albatross H3876.
. Hauerina fragilissima (H. B. Brady). X 40. Levuka, Fiji.
. Hauerina pacifica Cushman. X 40. Fig. 10, Guam Anchorage,
Ladrone Islands. Fig. 11, Mokaujar Anchorage, Fiji.
. Hauerina bradyi Cushman. X 55. Figs. 12, 18, Young micro-
spheric stages. Wavau Anchorge, Tonga Islands. Fig. 14,
Young megalospheric form. Levuka, Fiji. Fig. 15, Adult
specimen. Hereheretue Island.
Hauerina ornatissima (Karrer). X 40. Levuka, Fiji. Fig.
16, Adult. Fig. 17, Early stage.
Puate 11
Articulina lineata H. B. Brady. X 40. Fig. 1, Adult specimen.
Fig. 2, Early stage. a, Front view; 6, apertural view.
Mokaujar Anchorage, Fiji.
Articulina sulcata Reuss (?). > 40. Mokaujar Anchorage,
Fiji. Fig. 3, a, Front view; b, apertural view.
Nubeculina divaricata (H. B. Brady). X 40. Near Nairai,
Fiji.
Nubeculina chapmani, new species. XX 40. a, Front view; },
apertural view. Albatross H3898.
Nubeculina divaricata (H. B. Brady) var. advena Cushman.
X 40. a, Front view; b, apertural view. Mokaujar Anchor-
age, Fiji.
Sigmoilina edwardsi (Schlumberger). X 55. a, 6b, Opposite
sides; c, apertura] view. Albatross H3878.
Triloculina oblonga (Montagu). XX 40. a, 6, Opposite sides;
c,apertural view. Off Nairai, Fiji.
Triloculina circularis Bornemann. XX 75. a, b, Opposite sides;
c, apertural view. Vavau Anchorage, Tonga Islands.
Triloculina labiosa d’Orbigny. XX 75. a, b, Opposite sides; c,
apertural view. Rangiroa.
PuatTE 12
Triloculina labiosa d’Orbigny sparsicostata, new variety. X 565.
a, b, Opposite sides; c, apertural view. Off Hereheretue
Island.
Triloculina irregularis (d’Orbigny). X 40. a, 6b, Opposite
sides; c, apertural view. 40-50 fathoms, off Fiji.
TROPICAL PACIFIC FORAMINIFERA OF ALBATROSS 83
3 a-c.
4 a-c.
5 a-c.
6 a-c.
Figures la, b.
2 a, b.
3 a, b.
4 a,b.
6 a-c.
7 a-c.
Figures’ 1-9.
10, 11.
Fiaures 1-3.
4, 5.
6-8.
Triloculina oceanica, new species. XX 50. a,b, Opposite sides; c,
apertural view. Albatross H3935.
Triloculina oceanica flintiana, new species and variety. > 50.
a, b, Opposite sides; c, apertural view. Albatross H3857.
Triloculina spinata, new species. X 55. a, b, Opposite sides; c,
apertural view. Rongelap Atoll, Marshall Islands.
Triloculina fichteliana d’Orbigny. X 40. a, b, Opposite sides;
c, apertural view. Guam Anchorage, Ladrone Islands.
Puate 13
Triloculina trigonula (Lamarck) X 55. a, Front view; b,
apertural view. Mokaujar Anchorage, Fiji.
Triloculina austriaca d’Orbigny. XX 40. a, Front view; b,
apertural view. Rangiroa.
Triloculina tricarinata d’Orbigny. X 55. a, Front view; b,
apertural view. Viva Anchorage, Fiji.
Triloculina affinis d’Orbigny. X 55. a, Front view; b, aper-
tural view. Nairai, Fiji.
. Triloculina bertheliniana (H. B. Brady). X 55. Makemo La-
goon, Paumotu Islands.
Triloculina bicarinata d’Orbigny. X 40. a, b, Opposite sides; c,
apertural view. Mokaujar Anchorage, Fiji.
Triloculina sp. (?) XX 40. a, 6, Opposite sides; c, apertural
view. Rotonga.
PuatE 14
Pyrgo denticulata (H. B. Brady). X 40. Figs. 1-3, a, a, a,
Front views; b, b, 6b, apertural views. Mokaujar Anchorage,
Fiji. Figs. 4, 5, a, a, Front views; b, b, apertural views.
Guam Anchorage, Ladrone Islands. Fig. 6, a, Front view;
b, apertural view. Rongelap Atoll, Marshall Islands. Fig. 7,
a, Front view; b, apertural view. Albatross H3858. Figs.
8, 9, Levuka, Fiji.
Pyrgo denticulata (H. B. Brady) var. striolata (H. B. Brady).
x 40. Fig. 10, a, b, Opposite sides; c, apertural view.
Mokaujar Anchorage, Fiji.
Puate 15
Pyrgo murrhina (Schwager). X 40. a, a, a, Front views; b,
b, b, apertural views. Fig. 1, Albatross H8820. Fig. 2, Alba-
tross H3878. Fig. 3, Albatross H3841.
Pyrgo millettii (Cushman). XX 40. Fig. 4, a, Front view; 5,
apertural view. Levuka, Fiji.
Pyrgo globula (Bornemann). X 40. a, a, a, Front views; }, b, b,
apertural views. Figs. 6, 8, Albatross H3841. Fig. 7, Alba-
iross H8815.
84 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
PLatE 16
Fieures 1 a-c. Nodobacularia antillarum Cushman pacifica, new variety. a,
Front view; b, apertural view, X 40; c, early portion, X< 75.
Mokaujar Anchorage, Fiji.
2 a, b. Cornuspira involvens Reuss. XX 75. a, Front view; 6, periph-
eral view. Levuka, Fiji.
3, 4. Nodobacularia milletti, new species. Fig.3, Holotype. a, Front
view; b, apertural view, X 40; c, early portion, X 75. Fig.
4, Paratype. a, Front view, X 40; 6, early portion, X 75.
Mokaujar Anchorage, Fiji.
5 a, b. Spiropthalmidium acutimargo (H. B. Brady). X 75. a, Front
view; b, apertural view. Albatross H3829.
6 a-c. Planispirina auriculata Egger. X 75. a, b, Opposite sides; c,
apertural view. Port Lotten, Kersail, Caroline Islands.
7 a, b. Planispirina exigua (H. B. Brady). X 75. a, Front view;
b, apertural view. Levuka, Fiji.
8-10. Vertebralina striata d’Orbigny. X 40. Fig. 8, a, Front view;
b, apertural view. Fig. 10, Young specimen. Mokaujar
Anchorage, Fiji. Fig. 9, Coarsely ornamented form. Le-
vuka, Fiji.
PuatE 17
Fiegures 1-4. Parrina bradyi (Millett). X 55. Figs. 1, 2, 4, Hereheretue
Island. Fig. 3, 40-50 fathoms, off Fiji.
5 a-c. Fischerina helix Heron-Allen and Earland. X 75. a, Dorsal
view; 6b, ventral view; c, peripheral view. Hereheretue
Island.
6 a-c. Fischerina involuta, new species. XX 75. a, Dorsal view;
b, ventral view; c, peripheral view. Rangiroa.
7 a-c. Fischerina pellucida Millett. XX 75. a, Dorsal view; 6, ven-
tral view; c, peripheral view. Port Lotten, Kersail, Caroline
Islands.
8 a, b. Trochammina turbinata (H. B. Brady). X 75. a, Front view;
b, apertural view. Albatross H3973.
9 a,b. Nouria polymorphinoides Heron-Allen and Earland. X 30.
a. Front view: b, side view. Mokaujar Anchorage, Fiji.
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et eee
y
<<
et
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INDEX
abbreviata, Textularia, 11.
abyssorum, Rhabdammina, 4.
aceseata, Spiroloculina grateloupi, 35.
acutimargo, Spiroloculina, 41, 70.
Spiropthalmidium, 70.
Adelosina, 18.
striata, 18.
advena, Nubeculina divaricata, 49.
aequa, Spiroloculina antillarum, 38.
affinis, Triloculina, 58, 59.
affixa, Spiroloculina, 41.
agglutinans biformis, Textularia, 6.
agglutinans, Miliolina anguina, 18.
Quinqueloculina anguina, 18.
Textularia, 10.
agglutinatus, Reophax, 4.
albatrossi, Textularia, 12.
alveoliniformis, Massilina, 30.
Miliolina, 29.
Quinqueloculina, 29.
Schlumbergerina, 29.
Ammodiscidae, 5.
Ammodiscus incertus, 5.
anguina agglutinans, Miliolina, 18.
Quinqueloculina, 18.
angularis, Clavulina, 16, 17.
angularis difformis, Clavulina, 17.
angulata, Gaudryina triangularis, 14.
Spiroloculina antillarum, 36, 387.
Spiroloculina grata, 37.
angulosa, Spiroloculina, 34.
antillarum aequa, Spiroloculina, 38.
angulata, Spiroloculina, 36, 37.
pacifica, Nodobacularia, 68.
reticosa, Spiroloculina, 38.
antillarum, Nodobacularia, 69.
Spiroloculina, 36.
antillea reticosa, Spiroloculina, 38.
areniphora, Schlumbergerina, 29.
Articulina, 46.
conico-articulata, 68, 69.
lineata, 47.
nitida, 47.
suleata, 47.
asperula, Massilina, 31.
Spiroloculina, 31.
Astrorhizidae, 4.
auberiana, Quinqueloculina, 24.
auriculata, Planispirina, 72.
australis, Massilina, 32.
austriaca, Triloculina, 57.
bertheliniana, Miliolina, 60.
Miliolina tricarinata, 60.
Triloculina, 60.
berthelotiana, Quinqueloculina, 19.
bicarinata, Triloculina, 60, 61.
bidentata, Quinqueloculina, 20.
biformis, Textularia agglutinans, 6.
91513—32
Biloculina, 61.
bulloides, 61.
denticulata, 62.
denticulata striolata, 63.
depressa murrhyna, 64.
globulus, 65.
millettii, 66.
murrhina, 64.
ringens denticulata, 62.
ringens striolata, 63.
bradyi, Gaudryina, 138.
Hauerina, 44.
Nubecularia, 74.
Parrina, 74.
Silvestria, 74.
brongniartiana, Triloculina, 28.
bulloides, Biloculina, 61.
caduca, Spiroloculina, 39.
canaliculata, Spiroloculina, 35.
candeiana, Textularia, 9, 10, 11.
Textularia sagittula, 9.
chapmani, Nubeculina, 49.
circularis, Triloculina, 52.
clara, Spiroloculina, 40.
Clavulina, 16.
angularis, 16, 17.
angularis difformis, 17.
communis, 16.
difformis, 17.
pacifica, 16, 17.
parisiensis, 16.
tricarinata, 17.
communis, Clavulina, 16.
Planispirina, 71.
compressa, Hauerina, 42.
coneava, Textularia, 12, 13.
conica corrugata, Textularia, 12.
econica, Textularia, 11, 12.
conico-articulata, Articulina, 68, 69.
Cornuspira, 67, 71.
involvens, 67,
planorbis, 67.
Cornuspirinae, 67.
corrugata, Textularia, 12.
Textularia conica, 12.
costata, Miliolina, 21.
Quinqueloculina, 20.
crassa, Miliolina, 21.
Quinqueloculina, 21.
crassa subcuneata, Quinqueloculina, 21.
crenata, Massilina, 33.
Spiroloculina, 33.
erenulata, Quinqueloculina, 21.
cuvieriana, Quinqueloculina, 24.
denticulata, Biloculina, 62.
Biloculina ringens, 62.
Pyrgo, 62.
86 BULLETIN 161, UNITED STATES NATIONAL MUSEUM
denticulata striolata, Biloculina, 68.
striolata, Pyrgo, 63.
depressa murrhyna, Biloculina, 64.
depressa, Spiroloculina, 34, 40.
difformis, Clavulina, 17.
Clavulina angularis, 17.
divaricata advena, Nubeculina, 49.
divaricata, Nubecularia, 48, 49,
Nubeculina, 48.
durandii, Miliolina, 66.
edwardsi, Planispirina (Sigmoilina ), 45.
Sigmoilina, 45.
elongata, Pyrgo, 62.
excavata, Spiroloculina, 34.
exigua, Hauerina, 71.
Planispirina, 71.
eximia, Spiroloculina, 39.
\
fichteliana, Triloculina, 55.
Fischerina, 75.
helix, 76.
involuta, 76.
pellucida, 75.
rhodiensis, 75.
Fischerinidae, 75.
flintiana, Triloculina oceanica, 55.
Flintina, 37.
foliacea oceanica, Textularia, 8.
foliacea, Textularia, 8.
fragilissima, Hauerina, 42.
Spiroloculina, 42.
funafutiensis, Miliolina, 22.
Quinqueloculina, 22.
Gaudryina, 6, 13, 15.
bradyi, 13.
pupoides, 13.
quadrangularis, 14.
rugosa, 13.
rugulosa, 15.
sp.(?), 15.
triangularis angulata, 14.
Globotextulariinae, 77.
globula, Pyrgo, 65.
globulus, Biloculina, 65.
gracilis, Quinqueloculina Cri 23.
Grammostomum, 7.
grata angulata, Spiroloculina, 37.
reticosa, Spiroloculina, 38.
grata, Spiroloculina, 36, 37.
grateloupi acescata, Spiroloculina, 35
serrulata, Spiroloculina, 35.
grateloupi, Spiroloculina, 34, 36.
gualtieriana, Quinqueloculina, 23.
Haplophragmfum turbinatum, 77.
Haplophragmoides trullissata, 5.
Hauerina, 42, 43.
bradyi, 44.
compressa, 42.
exigua, 71.
fragilissima, 42.
ornatissima, 42, 43.
pacifiea, 44.
helix, Fischerina, 76.
impressa, Spiroloculina, 34.
inaequilateralis, Spiroloculina, 41.
inaequalis, Massilina, 32.
incertus, Ammodiscus, 5.
indica, Uniloculina, 18.
inflata, Nubecularia, 74.
involuta, Fischerina, 76.
involvens, Cornuspira, 67.
irregularis, Quinqueloculina, 54.
Triloculina, 54.
jonesi, Textularia, 7.
labiata, Textilaria, 7.
labiosa, Miliolina, 53.
Triloculina, 53.
labiosa sparsicostata, Triloculina, 54.
laevis, Pyrgo, 61.
lamarckiana, Quinqueloculina, 24.
lineata, Articulina, 47.
Lituolidae, 5.
macilenta, Massilina, 33.
Massilina secans, 33.
Miliolina, 33.
Quinqueloculina, 33.
Massilina, 23, 29, 30, 31, 34.
alveoliniformis, 30.
asperula, 31.
australis, 32.
crenata, 33.
inaequalis, 32.
macilenta, 33.
planata, 31.
secans, 32.
secans macilenta, 33.
mexicana, Textularia, 7.
Miliola, 34, 50, 61.
trigonula, 50, 56.
Miliolidae, 17.
Miliolina, 18, 29, 30, 50.
alveoliniformis, 29.
anguina agglutinans, 18.
bertheliniana, 60.
costata, 21.
crassa, 21.
durandii, 66.
funafutiensis, 22.
labiosa, 53.
macilenta, 33.
oblonga, 51.
parkeri, 25.
tricarinata, 59, 60.
tricarinata bertheliniana, 60.
trigonula, 56.
undulata, 27.
milletti, Nodobacularia, 69.
Spiroplectammina, 6, 7.
Textularia, 7.
millettii, Biloculina, 66.
Pyrgo, 66.
murrhina, Biloculina, 64.
Pyrgo, 64.
murrhyna, Biloculina depressa, 64.
Nautilus, 46.
nitida, Articulina, 47.
INDEX
Nodobacularia, 68.
antillarum, 69.
antillarum pacifica, 68.
milletti, 69.
Nodobaculariinae, 68.
Nouria, 77.
polymorphinoides, 77, 78.
Nubecularia, 48, 68, 74.
bradyi, 74.
divaricata, 48, 49.
inflata, 74.
tibia, 68.
Nubeculariinae, 74.
Nubeculina, 48.
chapmani, 49.
divaricata, 48.
divaricata advena, 49.
nussdorfensis, Quinqueloculina, 28.
oblonga, Miliolina, 51.
Triloculina, 50.
oblongum, Vermiculum, 50.
oceanica flintiana, Triloculina, 55.
oceanica, Textularia foliacea, 8.
Triloculina, 54.
Operculina, 67.
Ophthalmidiidae, 67.
Ophthalmidiinae, 70.
Ophthalmidium, 70.
Orbis, 67.
ornatissima, Hauerina, 42, 43.
Quinqueloculina, 48, 44.
pacifica, Clavulina, 16, 17.
Hauerina, 44.
Nodobacularia antillarum, 68.
Palaeotextularia, 7.
parisiensis, Clavulina, 16.
parkeri, Miliolina, 25.
Quinqueloculina, 25.
Parrina, 74.
bradyi, 74.
pellucida, Fischerina, 75.
planata, Massilina, 31.
Planispirina, 43, 45, 71.
auriculata, 72.
communis, 71.
exigua, 71.
sigmoidea, 45.
(Sigmoilina) edwardsi, 45.
planissima samoaensis, Spiroloculina,
39
planissima, Spiroloculina, 31.
planorbis, Cornuspira, 67.
Plecanium, 7.
polygona, Quinqueloculina, 25.
polymorphinoides, Nouria, 77, 78.
pupoides, Gaudryina, 13.
Pyrgo, 61.
denticulata, 62.
denticulata striolata, 63.
elongata, 62.
globula, 65.
laevis, 61.
millettii, 66.
murrhina, 6.
87
quadrangularis, Gaudryina, 14.
Quinqueloculina, 17, 25, 30, 45, 50, Gl.
alveoliniformis, 29.
anguina agglutinans, 18.
auberiana, 24.
berthelotiana, 19.
bidentata, 20.
eostata, 20.
crassa, 21.
crassa subcuneata, 21,
erenulata, 21.
cuvieriana, 24.
funafutiensis, 22.
ef. gracilis, 23.
gualteriana, 23.
irregularis, 54.
lamarckiana, 24.
macilenta, 33.
nussdorfensis, 28.
ornatissima, 48, 44.
parkeri, 25.
polygona, 25.
samoaensis, 26.
secans, 30.
semireticulosa, 27,
striatula, 27.
suleata, 28.
tricarinata, 22.
undulata, 28.
variabilis, 31.
Reophacidae, 4.
Reophax agglutinatus, 4.
reticosa, Spiroloculina antillarum, 38.
Spiroloculina antillea, 38.
Spiroloculina grata, 38.
reticulata, Triloculina, 61.
reticulosa, Vertebralina, 73.
Rhabdammina abyssorum, 4.
rhodiensis, Fischerina, 75.
ringens denticulata, Biloculina, 62.
striolata, Biloculina, 63.
rugosa, Gaudryina, 13.
Textularia, 15.
rugulosa, Gaudryina, 15.
sagittula candeiana, Textularia, 9.
sagittula, Textularia, 6, 7.
Sagrina, 48.
Samoaensis, Quinqueloculina, 26.
Spiroloculina planissima, 39.
Schlumbergerina, 29.
alveoliniformis, 29.
areniphora, 29.
secans macilenta, Massilina, 33.
secans, Massilina, 32.
Quinqueloculina, 30.
semialata, Textularia, 9.
seminulum, Serpula, 18.
semireticulosa, Quinqueloculina, 27.
Serpula, 18.
seminulum, 18.
serrulata, Spiroloculina grateloupi, 35.
sigmoidea, Planispirina, 45.
Sigmoilina, 45.
edwardsi, 45.
(Sigmoilina) edwardsi, Planispirina,
45,
88
Silvestria, 74.
bradyi, 74.
sparsicostata, Triloculina labiosa, 54.
spinata, Triloculina, 56.
Spirillina, 67.
Spiroloculina, 30, 31, 34, 45, 70.
acutimargo, 41, 70.
affixa, 41.
angulosa, 34.
antillarum, 36.
antillarum aequa, 38.
antillarum angulata, 36.
antillarum reticosa, 388.
antillea reticosa, 38.
asperula, 31.
ecaduca, 39.
eanaliculata, 35.
elara, 40.
crenata, 33.
depressa, 34, 40.
excavata, 34.
eximia, 389.
fragilissima, 42.
grata, 36, 37.
grata angulata, 87.
grata reticosa, 38.
grateloupi, 34, 36.
grateloupi acescata, 35.
grateloupi Serrulata, 35.
impressa, 34.
inaequilateralis, 41.
planissima, 31.
planissima samoaensis, 39.
sp.(?), 42.
striata, 39.
Spiroplecta, 6.
wrightii, 6.
Spiroplectammina, 6, 13.
milletti, 6, 7.
Spiroplectammininae, 6.
Spiropthalmidium, 70.
acutimargo, 70.
striata, Adelosina, 18.
Spiroloculina, 39.
Vertebralina, 72, 73.
striatula, Quinqueloculina, 27.
striolata, Biloculina denticulata, 63.
Biloculina ringens, 638.
Pyrgo denticulata, 63.
subcuneata, Quinqueloculina crassa, 21.
suborbicularis, Triloculina, 28.
sulcata, Articulina, 47.
Quinqueloculina, 28.
Textilaria, 7.
labiata, 7.
Textularia, 6, 7.
abbreviata, 11.
agglutinans, 10.
agglutinans biformis, 6.
albatrossi, 12.
candeiana, 9, 10, 11.
coneava, 12, 13.
conica, 11, 12.
conica corrugata, 12.
corrugata, 12.
O
BULLETIN 161, UNITED STATES NATIONAL MUSEUM
Textularia, foliacea, 8.
foliacea oceanica, 8.
jonesi, 7.
mexicana, 7.
milletti, 7.
rugosa, 15.
sagittula, 6, 7.
sagittula candeiana, 9.
semialata, 9.
Textulariidae, 6.
Textulariinae, 7.
tibia, Nubecularia, 68.
triangularis angulata, Gaudryina, 14.
tricarinata bertheliniana, Miliolina, 60.
tricarinata, Clavulina, 17.
Miliolina, 59, 60.
Quinqueloculina, 22.
Triloculina, 59.
trigonula, Miliola, 50, 56.
Miliolina, 56.
Triloculina, 56, 58.
Triloculina, 50, 61.
affinis, 58, 59.
austriaca, 57.
bertheliniana, 60.
bicarinata, 60, 61.
brongniartiana, 28.
circularis, 52.
fichteliana, 55.
irregularis, 54.
labiosa, 53.
labiosa sparsicostata, 54.
oblonga, 50.
oceanica, 54.
oceanica flintiana, 55.
reticulata, 61.
sp.(?), 61.
spinata, 56.
suborbicularis, 28.
tricarinata, 59.
trigonula, 56, 58.
Tritaxia, 16.
Trochammina trullissata, 5.
turbinata, 77.
Trochamminidae, 77.
Trochammininae, 77.
trullissata, Haplophragmoides, 5.
Trochammina, 5.
turbinata, Trochammina, 77.
turbinatum, Haplophragmium, 77.
undulata, Miliolina, 27.
Quinqueloculina, 28.
Uniloculina, 18.
indica, 18.
Valvulina, 16.
variabilis, Quinqueloculina, 31.
Vermiculum oblongum, 50.
Verneuilina, 16.
Verneuilinidae, 18.
Vertebralina, 46, 72.
reticulosa, 738.
striata, 72, 73.
wrightii, Spiroplecta, 6.
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